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Search by protein name, UniProt number, IPI number, or 15 AA P-site sequence.
Updated November 2019
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Protein Info
Short Name:
MUC5AC
Full Name:
Mucin-5AC
Alias:
Mucin-5 subtype AC, tracheobronchial
Type:
Mass (Da):
526590
Number AA:
5030
UniProt ID:
P98088
International Prot ID:
IPI00103397
Sequence:
Retrieve full protein sequence
GO Terms
External Links
Internal Links
Cellular Component:
GO:0005576
GO:0005578
GO:0031012
Uniprot
OncoNet
Molecular Function:
GO:0005198
GO:0005201
GO:0005488
PhosphoSite+
KinaseNET
Biological Process:
GO:0006915
GO:0006917
GO:0007155
Phosida
TranscriptoNet
STRING
Kinexus Products
Info Box
A variety of parameters were considered in the selection of putative P-sites. Confirmed P-Sites have lower Hydrophobicity Scores. The P-site Similarity Score is lower the more that it resembles typical confirmed corresponding P-Ser, P-Thr or P-Tyr sites. The Maximum KInase Score provides the calculated score for highest match of 500 human protein kinases for the amino acid sequence surrounding the target P-site as determined with Kinase Substrate Predictor V2. The Sum KInase Score provides the additive sum of the positive individual Kinase Substrate Predictor V2 scores from 500 human protein kinases. The Conservation Score is the average of the percent similarity of the human P-site with the equivalent P-site in 20 other diverse species. Click the coloured buttons below to retrieve this and other information in the Info Box, or click the orange buttons for relevant links.
Phosphosites
-7
-6
-5
-4
-3
-2
-1
0
1
2
3
4
5
6
7
Expt. conf.
Effect
Kinase
PPase
Kinexus Products
Ref.
Evol.
Kinase Pred.
P-site Match
Site 1
T24
A
L
A
C
T
R
H
T
G
H
A
Q
D
G
S
Site 2
S31
T
G
H
A
Q
D
G
S
S
E
S
S
Y
K
H
Site 3
S32
G
H
A
Q
D
G
S
S
E
S
S
Y
K
H
H
Site 4
S35
Q
D
G
S
S
E
S
S
Y
K
H
H
P
A
L
Site 5
Y36
D
G
S
S
E
S
S
Y
K
H
H
P
A
L
S
Site 6
S43
Y
K
H
H
P
A
L
S
P
I
A
R
G
P
S
Site 7
S50
S
P
I
A
R
G
P
S
G
V
P
L
R
G
A
Site 8
S72
T
I
P
V
V
R
A
S
N
P
A
H
N
G
R
Site 9
Y89
S
T
W
G
S
F
H
Y
K
T
F
D
G
D
V
Site 10
Y105
R
F
P
G
L
C
N
Y
V
F
S
E
H
C
G
Site 11
Y115
S
E
H
C
G
A
A
Y
E
D
F
N
I
Q
L
Site 12
S125
F
N
I
Q
L
R
R
S
Q
E
S
A
A
P
T
Site 13
S128
Q
L
R
R
S
Q
E
S
A
A
P
T
L
S
R
Site 14
T132
S
Q
E
S
A
A
P
T
L
S
R
V
L
M
K
Site 15
S173
S
G
V
L
I
Q
Q
S
S
S
Y
T
K
V
E
Site 16
S174
G
V
L
I
Q
Q
S
S
S
Y
T
K
V
E
A
Site 17
S175
V
L
I
Q
Q
S
S
S
Y
T
K
V
E
A
R
Site 18
T177
I
Q
Q
S
S
S
Y
T
K
V
E
A
R
L
G
Site 19
S194
L
M
W
N
H
D
D
S
L
L
L
E
L
D
T
Site 20
T201
S
L
L
L
E
L
D
T
K
Y
A
N
K
T
C
Site 21
Y203
L
L
E
L
D
T
K
Y
A
N
K
T
C
G
L
Site 22
S225
P
V
V
S
E
L
L
S
H
N
T
K
L
T
P
Site 23
T231
L
S
H
N
T
K
L
T
P
M
E
F
G
N
L
Site 24
T245
L
Q
K
M
D
D
P
T
E
Q
C
Q
D
P
V
Site 25
S260
P
E
P
P
R
N
C
S
T
G
F
G
I
C
E
Site 26
Y314
V
C
H
T
L
A
E
Y
S
R
Q
C
T
H
A
Site 27
T319
A
E
Y
S
R
Q
C
T
H
A
G
G
L
P
Q
Site 28
S356
S
P
C
A
D
T
C
S
N
Q
E
H
S
R
A
Site 29
S443
V
L
G
G
A
H
F
S
T
F
D
G
K
Q
Y
Site 30
T444
L
G
G
A
H
F
S
T
F
D
G
K
Q
Y
T
Site 31
Y450
S
T
F
D
G
K
Q
Y
T
V
H
G
D
C
S
Site 32
Y458
T
V
H
G
D
C
S
Y
V
L
T
K
P
C
D
Site 33
T461
G
D
C
S
Y
V
L
T
K
P
C
D
S
S
A
Site 34
T470
P
C
D
S
S
A
F
T
V
L
A
E
L
R
R
Site 35
T481
E
L
R
R
C
G
L
T
D
S
E
T
C
L
K
Site 36
S483
R
R
C
G
L
T
D
S
E
T
C
L
K
S
V
Site 37
S489
D
S
E
T
C
L
K
S
V
T
L
S
L
D
G
Site 38
T491
E
T
C
L
K
S
V
T
L
S
L
D
G
A
Q
Site 39
T567
A
P
K
L
R
G
Q
T
C
G
L
C
G
N
F
Site 40
S612
A
C
P
N
I
R
N
S
F
E
D
P
C
S
L
Site 41
S618
N
S
F
E
D
P
C
S
L
S
V
E
N
E
K
Site 42
S620
F
E
D
P
C
S
L
S
V
E
N
E
K
Y
A
Site 43
Y626
L
S
V
E
N
E
K
Y
A
Q
H
W
C
S
Q
Site 44
T702
G
V
C
T
K
P
M
T
T
C
P
K
S
M
T
Site 45
T703
V
C
T
K
P
M
T
T
C
P
K
S
M
T
Y
Site 46
Y712
P
K
S
M
T
Y
H
Y
H
V
S
T
C
Q
P
Site 47
S723
T
C
Q
P
T
C
R
S
L
S
E
G
D
I
T
Site 48
S725
Q
P
T
C
R
S
L
S
E
G
D
I
T
C
S
Site 49
T747
G
C
I
C
P
K
G
T
F
L
D
D
T
G
K
Site 50
T752
K
G
T
F
L
D
D
T
G
K
C
V
Q
A
S
Site 51
S775
S
M
I
P
N
G
E
S
V
H
D
S
G
A
I
Site 52
T815
F
F
D
C
R
N
A
T
P
G
D
T
G
A
G
Site 53
Y835
H
T
L
D
M
T
C
Y
S
P
Q
C
V
P
G
Site 54
T877
A
S
Y
R
A
G
Q
T
I
R
V
G
C
N
T
Site 55
Y911
A
V
Y
G
D
G
H
Y
L
T
F
D
G
Q
S
Site 56
T913
Y
G
D
G
H
Y
L
T
F
D
G
Q
S
Y
S
Site 57
Y927
S
F
N
G
D
C
E
Y
T
L
V
Q
N
H
C
Site 58
S939
N
H
C
G
G
K
D
S
T
Q
D
S
F
R
V
Site 59
T940
H
C
G
G
K
D
S
T
Q
D
S
F
R
V
V
Site 60
S943
G
K
D
S
T
Q
D
S
F
R
V
V
T
E
N
Site 61
T948
Q
D
S
F
R
V
V
T
E
N
V
P
C
G
T
Site 62
T985
G
K
V
E
V
I
G
T
D
E
S
Q
E
V
P
Site 63
S988
E
V
I
G
T
D
E
S
Q
E
V
P
Y
T
I
Site 64
T994
E
S
Q
E
V
P
Y
T
I
Q
Q
M
G
I
Y
Site 65
S1053
D
F
A
T
R
S
R
S
V
V
G
D
V
L
E
Site 66
S1064
D
V
L
E
F
G
N
S
W
K
L
S
P
S
C
Site 67
S1068
F
G
N
S
W
K
L
S
P
S
C
P
D
A
L
Site 68
S1070
N
S
W
K
L
S
P
S
C
P
D
A
L
A
P
Site 69
T1082
L
A
P
K
D
P
C
T
A
N
P
F
R
K
S
Site 70
Y1115
A
H
V
E
P
A
R
Y
Y
E
A
C
V
N
D
Site 71
Y1116
H
V
E
P
A
R
Y
Y
E
A
C
V
N
D
A
Site 72
Y1207
V
R
G
L
E
G
C
Y
P
K
C
P
P
E
A
Site 73
T1230
Q
C
V
A
T
C
P
T
P
P
L
P
P
R
C
Site 74
S1243
R
C
H
V
H
G
K
S
Y
R
P
G
A
V
V
Site 75
T1269
T
E
R
G
V
E
C
T
Y
K
A
E
A
C
V
Site 76
Y1279
A
E
A
C
V
C
T
Y
N
G
Q
R
F
H
P
Site 77
Y1291
F
H
P
G
D
V
I
Y
H
T
T
D
G
T
G
Site 78
T1293
P
G
D
V
I
Y
H
T
T
D
G
T
G
G
C
Site 79
T1297
I
Y
H
T
T
D
G
T
G
G
C
I
S
A
R
Site 80
S1302
D
G
T
G
G
C
I
S
A
R
C
G
A
N
G
Site 81
Y1316
G
T
I
E
R
R
V
Y
P
C
S
P
T
T
P
Site 82
S1319
E
R
R
V
Y
P
C
S
P
T
T
P
V
P
P
Site 83
T1321
R
V
Y
P
C
S
P
T
T
P
V
P
P
T
T
Site 84
T1322
V
Y
P
C
S
P
T
T
P
V
P
P
T
T
F
Site 85
T1327
P
T
T
P
V
P
P
T
T
F
S
F
S
T
P
Site 86
T1328
T
T
P
V
P
P
T
T
F
S
F
S
T
P
P
Site 87
S1339
S
T
P
P
L
V
V
S
S
T
H
T
P
S
N
Site 88
S1340
T
P
P
L
V
V
S
S
T
H
T
P
S
N
G
Site 89
T1341
P
P
L
V
V
S
S
T
H
T
P
S
N
G
P
Site 90
T1343
L
V
V
S
S
T
H
T
P
S
N
G
P
S
S
Site 91
S1345
V
S
S
T
H
T
P
S
N
G
P
S
S
A
H
Site 92
S1349
H
T
P
S
N
G
P
S
S
A
H
T
G
P
P
Site 93
S1350
T
P
S
N
G
P
S
S
A
H
T
G
P
P
S
Site 94
T1353
N
G
P
S
S
A
H
T
G
P
P
S
S
A
W
Site 95
S1357
S
A
H
T
G
P
P
S
S
A
W
P
T
T
A
Site 96
S1358
A
H
T
G
P
P
S
S
A
W
P
T
T
A
G
Site 97
T1362
P
P
S
S
A
W
P
T
T
A
G
T
S
P
R
Site 98
T1363
P
S
S
A
W
P
T
T
A
G
T
S
P
R
T
Site 99
S1367
W
P
T
T
A
G
T
S
P
R
T
R
L
P
T
Site 100
T1370
T
A
G
T
S
P
R
T
R
L
P
T
A
S
A
Site 101
T1374
S
P
R
T
R
L
P
T
A
S
A
S
L
P
P
Site 102
S1376
R
T
R
L
P
T
A
S
A
S
L
P
P
V
C
Site 103
T1402
V
S
R
P
G
R
G
T
D
S
G
D
F
D
T
Site 104
S1404
R
P
G
R
G
T
D
S
G
D
F
D
T
L
E
Site 105
T1409
T
D
S
G
D
F
D
T
L
E
N
L
R
A
H
Site 106
Y1418
E
N
L
R
A
H
G
Y
R
V
C
E
S
P
R
Site 107
S1423
H
G
Y
R
V
C
E
S
P
R
S
V
E
C
R
Site 108
S1426
R
V
C
E
S
P
R
S
V
E
C
R
A
E
D
Site 109
T1455
C
S
P
D
V
G
L
T
C
R
N
R
E
Q
A
Site 110
Y1467
E
Q
A
S
G
L
C
Y
N
Y
Q
I
R
V
Q
Site 111
S1482
C
C
T
P
L
P
C
S
T
S
S
S
P
A
Q
Site 112
S1484
T
P
L
P
C
S
T
S
S
S
P
A
Q
T
T
Site 113
S1485
P
L
P
C
S
T
S
S
S
P
A
Q
T
T
P
Site 114
S1486
L
P
C
S
T
S
S
S
P
A
Q
T
T
P
P
Site 115
T1490
T
S
S
S
P
A
Q
T
T
P
P
T
T
S
K
Site 116
T1491
S
S
S
P
A
Q
T
T
P
P
T
T
S
K
T
Site 117
T1494
P
A
Q
T
T
P
P
T
T
S
K
T
T
E
T
Site 118
S1504
K
T
T
E
T
R
A
S
G
S
S
A
P
S
S
Site 119
S1506
T
E
T
R
A
S
G
S
S
A
P
S
S
T
P
Site 120
S1507
E
T
R
A
S
G
S
S
A
P
S
S
T
P
G
Site 121
S1510
A
S
G
S
S
A
P
S
S
T
P
G
T
V
S
Site 122
S1511
S
G
S
S
A
P
S
S
T
P
G
T
V
S
L
Site 123
T1512
G
S
S
A
P
S
S
T
P
G
T
V
S
L
S
Site 124
T1515
A
P
S
S
T
P
G
T
V
S
L
S
T
A
R
Site 125
S1517
S
S
T
P
G
T
V
S
L
S
T
A
R
T
T
Site 126
S1519
T
P
G
T
V
S
L
S
T
A
R
T
T
P
A
Site 127
T1520
P
G
T
V
S
L
S
T
A
R
T
T
P
A
P
Site 128
T1523
V
S
L
S
T
A
R
T
T
P
A
P
G
T
A
Site 129
T1524
S
L
S
T
A
R
T
T
P
A
P
G
T
A
T
Site 130
S1532
P
A
P
G
T
A
T
S
V
K
K
T
F
S
T
Site 131
T1536
T
A
T
S
V
K
K
T
F
S
T
P
S
P
P
Site 132
S1538
T
S
V
K
K
T
F
S
T
P
S
P
P
P
V
Site 133
T1539
S
V
K
K
T
F
S
T
P
S
P
P
P
V
P
Site 134
S1541
K
K
T
F
S
T
P
S
P
P
P
V
P
A
T
Site 135
T1548
S
P
P
P
V
P
A
T
S
T
S
S
M
S
T
Site 136
S1549
P
P
P
V
P
A
T
S
T
S
S
M
S
T
T
Site 137
T1550
P
P
V
P
A
T
S
T
S
S
M
S
T
T
A
Site 138
S1551
P
V
P
A
T
S
T
S
S
M
S
T
T
A
P
Site 139
S1554
A
T
S
T
S
S
M
S
T
T
A
P
G
T
S
Site 140
T1555
T
S
T
S
S
M
S
T
T
A
P
G
T
S
V
Site 141
T1556
S
T
S
S
M
S
T
T
A
P
G
T
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Site 445
T3149
T
T
T
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S
A
P
T
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S
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T
S
A
P
Site 446
S3150
T
T
S
S
A
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T
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S
T
T
S
A
P
T
Site 447
S3151
T
S
S
A
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S
S
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T
S
A
P
T
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Site 448
T3152
S
S
A
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S
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T
S
A
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T
T
S
Site 449
T3153
S
A
P
T
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S
T
T
S
A
P
T
T
S
T
Site 450
S3154
A
P
T
S
S
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T
S
A
P
T
T
S
T
I
Site 451
T3158
S
T
T
S
A
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T
T
S
T
I
S
A
P
T
Site 452
S3159
T
T
S
A
P
T
T
S
T
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S
A
P
T
T
Site 453
T3166
S
T
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S
A
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T
S
T
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S
A
P
T
Site 454
S3167
T
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A
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T
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T
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S
A
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T
T
Site 455
S3175
T
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S
A
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T
S
T
T
S
A
P
T
A
Site 456
T3177
S
A
P
T
T
S
T
T
S
A
P
T
A
S
T
Site 457
S3183
T
T
S
A
P
T
A
S
T
T
S
A
P
T
S
Site 458
T3185
S
A
P
T
A
S
T
T
S
A
P
T
S
T
S
Site 459
S3186
A
P
T
A
S
T
T
S
A
P
T
S
T
S
S
Site 460
S3190
S
T
T
S
A
P
T
S
T
S
S
A
P
T
T
Site 461
S3192
T
S
A
P
T
S
T
S
S
A
P
T
T
N
T
Site 462
T3196
T
S
T
S
S
A
P
T
T
N
T
T
S
A
P
Site 463
T3197
S
T
S
S
A
P
T
T
N
T
T
S
A
P
T
Site 464
T3200
S
A
P
T
T
N
T
T
S
A
P
T
T
S
T
Site 465
T3205
N
T
T
S
A
P
T
T
S
T
T
S
A
P
I
Site 466
S3206
T
T
S
A
P
T
T
S
T
T
S
A
P
I
T
Site 467
T3208
S
A
P
T
T
S
T
T
S
A
P
I
T
S
T
Site 468
T3213
S
T
T
S
A
P
I
T
S
T
I
S
A
P
T
Site 469
S3214
T
T
S
A
P
I
T
S
T
I
S
A
P
T
T
Site 470
T3221
S
T
I
S
A
P
T
T
S
T
T
S
T
P
Q
Site 471
S3222
T
I
S
A
P
T
T
S
T
T
S
T
P
Q
T
Site 472
T3224
S
A
P
T
T
S
T
T
S
T
P
Q
T
S
T
Site 473
T3226
P
T
T
S
T
T
S
T
P
Q
T
S
T
I
S
Site 474
T3229
S
T
T
S
T
P
Q
T
S
T
I
S
S
P
T
Site 475
S3230
T
T
S
T
P
Q
T
S
T
I
S
S
P
T
T
Site 476
S3234
P
Q
T
S
T
I
S
S
P
T
T
S
T
T
P
Site 477
T3237
S
T
I
S
S
P
T
T
S
T
T
P
T
P
Q
Site 478
S3238
T
I
S
S
P
T
T
S
T
T
P
T
P
Q
T
Site 479
T3240
S
S
P
T
T
S
T
T
P
T
P
Q
T
S
T
Site 480
T3242
P
T
T
S
T
T
P
T
P
Q
T
S
T
T
S
Site 481
T3245
S
T
T
P
T
P
Q
T
S
T
T
S
S
P
T
Site 482
S3246
T
T
P
T
P
Q
T
S
T
T
S
S
P
T
T
Site 483
T3248
P
T
P
Q
T
S
T
T
S
S
P
T
T
S
T
Site 484
S3250
P
Q
T
S
T
T
S
S
P
T
T
S
T
T
S
Site 485
T3253
S
T
T
S
S
P
T
T
S
T
T
S
A
P
T
Site 486
S3254
T
T
S
S
P
T
T
S
T
T
S
A
P
T
T
Site 487
T3256
S
S
P
T
T
S
T
T
S
A
P
T
T
S
T
Site 488
S3286
I
S
S
A
P
T
S
S
T
T
S
A
P
T
A
Site 489
T3287
S
S
A
P
T
S
S
T
T
S
A
P
T
A
S
Site 490
S3289
A
P
T
S
S
T
T
S
A
P
T
A
S
T
I
Site 491
S3294
T
T
S
A
P
T
A
S
T
I
S
A
P
T
T
Site 492
T3301
S
T
I
S
A
P
T
T
S
T
T
S
F
H
T
Site 493
S3302
T
I
S
A
P
T
T
S
T
T
S
F
H
T
T
Site 494
S3305
A
P
T
T
S
T
T
S
F
H
T
T
S
T
T
Site 495
T3309
S
T
T
S
F
H
T
T
S
T
T
S
P
P
T
Site 496
S3310
T
T
S
F
H
T
T
S
T
T
S
P
P
T
S
Site 497
T3311
T
S
F
H
T
T
S
T
T
S
P
P
T
S
S
Site 498
T3312
S
F
H
T
T
S
T
T
S
P
P
T
S
S
T
Site 499
S3313
F
H
T
T
S
T
T
S
P
P
T
S
S
T
S
Site 500
T3316
T
S
T
T
S
P
P
T
S
S
T
S
S
T
P
Site 501
S3317
S
T
T
S
P
P
T
S
S
T
S
S
T
P
Q
Site 502
S3318
T
T
S
P
P
T
S
S
T
S
S
T
P
Q
T
Site 503
S3320
S
P
P
T
S
S
T
S
S
T
P
Q
T
S
K
Site 504
S3321
P
P
T
S
S
T
S
S
T
P
Q
T
S
K
T
Site 505
T3322
P
T
S
S
T
S
S
T
P
Q
T
S
K
T
S
Site 506
T3325
S
T
S
S
T
P
Q
T
S
K
T
S
A
A
T
Site 507
S3333
S
K
T
S
A
A
T
S
S
T
T
S
G
S
G
Site 508
S3334
K
T
S
A
A
T
S
S
T
T
S
G
S
G
T
Site 509
T3335
T
S
A
A
T
S
S
T
T
S
G
S
G
T
T
Site 510
S3337
A
A
T
S
S
T
T
S
G
S
G
T
T
P
S
Site 511
S3339
T
S
S
T
T
S
G
S
G
T
T
P
S
P
V
Site 512
T3341
S
T
T
S
G
S
G
T
T
P
S
P
V
P
T
Site 513
T3342
T
T
S
G
S
G
T
T
P
S
P
V
P
T
T
Site 514
S3344
S
G
S
G
T
T
P
S
P
V
P
T
T
S
T
Site 515
T3535
S
T
T
S
A
P
T
T
S
T
I
P
A
S
T
Site 516
S3536
T
T
S
A
P
T
T
S
T
I
P
A
S
T
P
Site 517
T3537
T
S
A
P
T
T
S
T
I
P
A
S
T
P
S
Site 518
S3541
T
T
S
T
I
P
A
S
T
P
S
T
T
S
A
Site 519
T3542
T
S
T
I
P
A
S
T
P
S
T
T
S
A
P
Site 520
S3544
T
I
P
A
S
T
P
S
T
T
S
A
P
T
T
Site 521
T3545
I
P
A
S
T
P
S
T
T
S
A
P
T
T
S
Site 522
T3546
P
A
S
T
P
S
T
T
S
A
P
T
T
S
T
Site 523
S3547
A
S
T
P
S
T
T
S
A
P
T
T
S
T
T
Site 524
S3560
T
T
S
A
P
T
T
S
T
T
S
A
P
T
H
Site 525
T3562
S
A
P
T
T
S
T
T
S
A
P
T
H
R
T
Site 526
T3570
S
A
P
T
H
R
T
T
S
G
P
T
T
S
T
Site 527
S3571
A
P
T
H
R
T
T
S
G
P
T
T
S
T
T
Site 528
T3575
R
T
T
S
G
P
T
T
S
T
T
L
A
P
T
Site 529
S3576
T
T
S
G
P
T
T
S
T
T
L
A
P
T
T
Site 530
T3578
S
G
P
T
T
S
T
T
L
A
P
T
T
S
T
Site 531
T3583
S
T
T
L
A
P
T
T
S
T
T
S
A
P
T
Site 532
S3584
T
T
L
A
P
T
T
S
T
T
S
A
P
T
T
Site 533
T3586
L
A
P
T
T
S
T
T
S
A
P
T
T
S
T
Site 534
T3591
S
T
T
S
A
P
T
T
S
T
N
S
A
P
T
Site 535
S3592
T
T
S
A
P
T
T
S
T
N
S
A
P
T
T
Site 536
T3599
S
T
N
S
A
P
T
T
S
T
I
S
A
S
T
Site 537
T3601
N
S
A
P
T
T
S
T
I
S
A
S
T
T
S
Site 538
S3603
A
P
T
T
S
T
I
S
A
S
T
T
S
T
I
Site 539
S3605
T
T
S
T
I
S
A
S
T
T
S
T
I
S
A
Site 540
T3607
S
T
I
S
A
S
T
T
S
T
I
S
A
P
T
Site 541
S3608
T
I
S
A
S
T
T
S
T
I
S
A
P
T
T
Site 542
T3609
I
S
A
S
T
T
S
T
I
S
A
P
T
T
S
Site 543
S3611
A
S
T
T
S
T
I
S
A
P
T
T
S
T
I
Site 544
T3615
S
T
I
S
A
P
T
T
S
T
I
S
S
P
T
Site 545
S3616
T
I
S
A
P
T
T
S
T
I
S
S
P
T
S
Site 546
T3617
I
S
A
P
T
T
S
T
I
S
S
P
T
S
S
Site 547
S3619
A
P
T
T
S
T
I
S
S
P
T
S
S
T
T
Site 548
S3620
P
T
T
S
T
I
S
S
P
T
S
S
T
T
S
Site 549
S3623
S
T
I
S
S
P
T
S
S
T
T
S
T
P
Q
Site 550
S3624
T
I
S
S
P
T
S
S
T
T
S
T
P
Q
T
Site 551
T3626
S
S
P
T
S
S
T
T
S
T
P
Q
T
S
K
Site 552
S3627
S
P
T
S
S
T
T
S
T
P
Q
T
S
K
T
Site 553
T3654
T
T
P
S
P
V
P
T
T
S
T
T
S
A
S
Site 554
T3655
T
P
S
P
V
P
T
T
S
T
T
S
A
S
T
Site 555
S3656
P
S
P
V
P
T
T
S
T
T
S
A
S
T
T
Site 556
T3657
S
P
V
P
T
T
S
T
T
S
A
S
T
T
S
Site 557
S3661
T
T
S
T
T
S
A
S
T
T
S
T
T
S
A
Site 558
T3663
S
T
T
S
A
S
T
T
S
T
T
S
A
P
T
Site 559
T3665
T
S
A
S
T
T
S
T
T
S
A
P
T
T
S
Site 560
T3666
S
A
S
T
T
S
T
T
S
A
P
T
T
S
T
Site 561
S3667
A
S
T
T
S
T
T
S
A
P
T
T
S
T
T
Site 562
T3671
S
T
T
S
A
P
T
T
S
T
T
S
G
P
G
Site 563
S3672
T
T
S
A
P
T
T
S
T
T
S
G
P
G
T
Site 564
T3674
S
A
P
T
T
S
T
T
S
G
P
G
T
T
P
Site 565
S3675
A
P
T
T
S
T
T
S
G
P
G
T
T
P
S
Site 566
T3679
S
T
T
S
G
P
G
T
T
P
S
P
V
P
S
Site 567
T3680
T
T
S
G
P
G
T
T
P
S
P
V
P
S
T
Site 568
S3682
S
G
P
G
T
T
P
S
P
V
P
S
T
S
I
Site 569
S3686
T
T
P
S
P
V
P
S
T
S
I
T
S
A
A
Site 570
T3687
T
P
S
P
V
P
S
T
S
I
T
S
A
A
T
Site 571
S3688
P
S
P
V
P
S
T
S
I
T
S
A
A
T
T
Site 572
T3690
P
V
P
S
T
S
I
T
S
A
A
T
T
S
T
Site 573
T3695
S
I
T
S
A
A
T
T
S
T
T
S
A
P
T
Site 574
S3696
I
T
S
A
A
T
T
S
T
T
S
A
P
T
T
Site 575
T3703
S
T
T
S
A
P
T
T
R
T
T
S
A
P
T
Site 576
T3706
S
A
P
T
T
R
T
T
S
A
P
T
S
S
M
Site 577
S3707
A
P
T
T
R
T
T
S
A
P
T
S
S
M
T
Site 578
S3711
R
T
T
S
A
P
T
S
S
M
T
S
G
P
G
Site 579
S3712
T
T
S
A
P
T
S
S
M
T
S
G
P
G
T
Site 580
T3714
S
A
P
T
S
S
M
T
S
G
P
G
T
T
P
Site 581
S3715
A
P
T
S
S
M
T
S
G
P
G
T
T
P
S
Site 582
T3719
S
M
T
S
G
P
G
T
T
P
S
P
V
P
T
Site 583
T3720
M
T
S
G
P
G
T
T
P
S
P
V
P
T
T
Site 584
S3752
P
S
P
V
P
T
T
S
T
T
S
A
P
I
T
Site 585
T3754
P
V
P
T
T
S
T
T
S
A
P
I
T
S
T
Site 586
T3759
S
T
T
S
A
P
I
T
S
T
T
S
G
P
G
Site 587
S3760
T
T
S
A
P
I
T
S
T
T
S
G
P
G
S
Site 588
T3761
T
S
A
P
I
T
S
T
T
S
G
P
G
S
T
Site 589
T3762
S
A
P
I
T
S
T
T
S
G
P
G
S
T
P
Site 590
S3763
A
P
I
T
S
T
T
S
G
P
G
S
T
P
S
Site 591
S3767
S
T
T
S
G
P
G
S
T
P
S
P
V
P
T
Site 592
T3768
T
T
S
G
P
G
S
T
P
S
P
V
P
T
T
Site 593
S3770
S
G
P
G
S
T
P
S
P
V
P
T
T
S
T
Site 594
T3774
S
T
P
S
P
V
P
T
T
S
T
T
S
A
P
Site 595
S3789
T
T
S
T
T
S
A
S
T
A
S
T
T
S
G
Site 596
S3792
T
T
S
A
S
T
A
S
T
T
S
G
P
T
T
Site 597
T3793
T
S
A
S
T
A
S
T
T
S
G
P
T
T
S
Site 598
T3794
S
A
S
T
A
S
T
T
S
G
P
T
T
S
T
Site 599
S3795
A
S
T
A
S
T
T
S
G
P
T
T
S
T
T
Site 600
T3799
S
T
T
S
G
P
T
T
S
T
T
S
A
S
T
Site 601
S3800
T
T
S
G
P
T
T
S
T
T
S
A
S
T
T
Site 602
T3801
T
S
G
P
T
T
S
T
T
S
A
S
T
T
S
Site 603
T3802
S
G
P
T
T
S
T
T
S
A
S
T
T
S
T
Site 604
S3803
G
P
T
T
S
T
T
S
A
S
T
T
S
T
I
Site 605
S3805
T
T
S
T
T
S
A
S
T
T
S
T
I
S
P
Site 606
T3807
S
T
T
S
A
S
T
T
S
T
I
S
P
L
T
Site 607
T3809
T
S
A
S
T
T
S
T
I
S
P
L
T
T
S
Site 608
S3811
A
S
T
T
S
T
I
S
P
L
T
T
S
T
T
Site 609
T3814
T
S
T
I
S
P
L
T
T
S
T
T
S
A
P
Site 610
T3818
S
P
L
T
T
S
T
T
S
A
P
I
T
S
M
Site 611
T3823
S
T
T
S
A
P
I
T
S
M
P
S
G
P
G
Site 612
S3824
T
T
S
A
P
I
T
S
M
P
S
G
P
G
T
Site 613
S3827
A
P
I
T
S
M
P
S
G
P
G
T
T
P
S
Site 614
T3831
S
M
P
S
G
P
G
T
T
P
S
P
V
P
T
Site 615
T3832
M
P
S
G
P
G
T
T
P
S
P
V
P
T
T
Site 616
S3880
T
T
S
A
S
T
A
S
T
T
S
G
P
G
T
Site 617
T3881
T
S
A
S
T
A
S
T
T
S
G
P
G
T
T
Site 618
S3883
A
S
T
A
S
T
T
S
G
P
G
T
T
P
S
Site 619
T3913
T
S
A
S
T
A
S
T
T
S
G
P
G
T
S
Site 620
T3914
S
A
S
T
A
S
T
T
S
G
P
G
T
S
L
Site 621
S3915
A
S
T
A
S
T
T
S
G
P
G
T
S
L
S
Site 622
T3919
S
T
T
S
G
P
G
T
S
L
S
P
V
P
T
Site 623
S3920
T
T
S
G
P
G
T
S
L
S
P
V
P
T
T
Site 624
S3922
S
G
P
G
T
S
L
S
P
V
P
T
T
S
T
Site 625
T3926
T
S
L
S
P
V
P
T
T
S
T
T
S
A
P
Site 626
T3927
S
L
S
P
V
P
T
T
S
T
T
S
A
P
T
Site 627
S3928
L
S
P
V
P
T
T
S
T
T
S
A
P
T
T
Site 628
T3974
T
T
P
S
P
V
P
T
T
S
T
T
P
V
S
Site 629
S3976
P
S
P
V
P
T
T
S
T
T
P
V
S
K
T
Site 630
T3977
S
P
V
P
T
T
S
T
T
P
V
S
K
T
S
Site 631
T3978
P
V
P
T
T
S
T
T
P
V
S
K
T
S
T
Site 632
S3981
T
T
S
T
T
P
V
S
K
T
S
T
S
H
L
Site 633
T3983
S
T
T
P
V
S
K
T
S
T
S
H
L
S
V
Site 634
S3984
T
T
P
V
S
K
T
S
T
S
H
L
S
V
S
Site 635
T3985
T
P
V
S
K
T
S
T
S
H
L
S
V
S
K
Site 636
S3986
P
V
S
K
T
S
T
S
H
L
S
V
S
K
T
Site 637
S3989
K
T
S
T
S
H
L
S
V
S
K
T
T
H
S
Site 638
S3991
S
T
S
H
L
S
V
S
K
T
T
H
S
Q
P
Site 639
T3993
S
H
L
S
V
S
K
T
T
H
S
Q
P
V
T
Site 640
S3996
S
V
S
K
T
T
H
S
Q
P
V
T
S
D
C
Site 641
T4000
T
T
H
S
Q
P
V
T
S
D
C
H
P
L
C
Site 642
S4056
R
L
Q
C
R
A
E
S
H
P
E
V
N
I
E
Site 643
T4110
T
P
R
G
C
P
V
T
S
V
T
P
Y
G
T
Site 644
S4111
P
R
G
C
P
V
T
S
V
T
P
Y
G
T
S
Site 645
T4113
G
C
P
V
T
S
V
T
P
Y
G
T
S
P
T
Site 646
Y4115
P
V
T
S
V
T
P
Y
G
T
S
P
T
N
A
Site 647
S4118
S
V
T
P
Y
G
T
S
P
T
N
A
L
Y
P
Site 648
Y4124
T
S
P
T
N
A
L
Y
P
S
L
S
T
S
M
Site 649
S4126
P
T
N
A
L
Y
P
S
L
S
T
S
M
V
S
Site 650
S4149
A
S
S
S
V
A
S
S
S
V
A
Y
S
T
Q
Site 651
S4150
S
S
S
V
A
S
S
S
V
A
Y
S
T
Q
T
Site 652
S4154
A
S
S
S
V
A
Y
S
T
Q
T
C
F
C
N
Site 653
Y4167
C
N
V
A
D
R
L
Y
P
A
G
S
T
I
Y
Site 654
S4171
D
R
L
Y
P
A
G
S
T
I
Y
R
H
R
D
Site 655
Y4184
R
D
L
A
G
H
C
Y
Y
A
L
C
S
Q
D
Site 656
S4200
Q
V
V
R
G
V
D
S
D
C
P
S
T
T
L
Site 657
S4204
G
V
D
S
D
C
P
S
T
T
L
P
P
A
P
Site 658
T4205
V
D
S
D
C
P
S
T
T
L
P
P
A
P
A
Site 659
T4206
D
S
D
C
P
S
T
T
L
P
P
A
P
A
T
Site 660
T4213
T
L
P
P
A
P
A
T
S
P
S
I
S
T
S
Site 661
S4214
L
P
P
A
P
A
T
S
P
S
I
S
T
S
E
Site 662
S4216
P
A
P
A
T
S
P
S
I
S
T
S
E
P
V
Site 663
S4218
P
A
T
S
P
S
I
S
T
S
E
P
V
T
E
Site 664
S4220
T
S
P
S
I
S
T
S
E
P
V
T
E
L
G
Site 665
T4224
I
S
T
S
E
P
V
T
E
L
G
C
P
N
A
Site 666
T4240
P
P
R
K
K
G
E
T
W
A
T
P
N
C
S
Site 667
T4243
K
K
G
E
T
W
A
T
P
N
C
S
E
A
T
Site 668
S4258
C
E
G
N
N
V
I
S
L
S
P
R
T
C
P
Site 669
S4260
G
N
N
V
I
S
L
S
P
R
T
C
P
R
V
Site 670
T4271
C
P
R
V
E
K
P
T
C
A
N
G
Y
P
A
Site 671
Y4276
K
P
T
C
A
N
G
Y
P
A
V
K
V
A
D
Site 672
Y4305
S
G
W
G
D
P
H
Y
I
T
F
D
G
T
Y
Site 673
Y4312
Y
I
T
F
D
G
T
Y
Y
T
F
L
D
N
C
Site 674
S4354
D
G
L
S
C
P
R
S
I
I
L
E
Y
H
Q
Site 675
T4367
H
Q
D
R
V
V
L
T
R
K
P
V
H
G
V
Site 676
S4387
I
F
N
N
K
V
V
S
P
G
F
R
K
N
G
Site 677
S4398
R
K
N
G
I
V
V
S
R
I
G
V
K
M
Y
Site 678
S4429
F
S
V
E
V
P
F
S
K
F
A
N
N
T
E
Site 679
T4443
E
G
Q
C
G
T
C
T
N
D
R
K
D
E
C
Site 680
T4452
D
R
K
D
E
C
R
T
P
R
G
T
V
V
A
Site 681
T4456
E
C
R
T
P
R
G
T
V
V
A
S
C
S
E
Site 682
T4484
A
C
H
R
P
H
P
T
P
T
T
V
G
P
T
Site 683
T4487
R
P
H
P
T
P
T
T
V
G
P
T
T
V
G
Site 684
T4507
P
T
T
V
G
S
T
T
V
G
P
T
T
P
P
Site 685
T4512
S
T
T
V
G
P
T
T
P
P
A
P
C
L
P
Site 686
Y4598
T
C
P
A
D
K
V
Y
Q
P
C
G
P
S
N
Site 687
S4607
P
C
G
P
S
N
P
S
Y
C
Y
G
N
D
S
Site 688
Y4608
C
G
P
S
N
P
S
Y
C
Y
G
N
D
S
A
Site 689
Y4610
P
S
N
P
S
Y
C
Y
G
N
D
S
A
S
L
Site 690
S4614
S
Y
C
Y
G
N
D
S
A
S
L
G
A
L
R
Site 691
S4616
C
Y
G
N
D
S
A
S
L
G
A
L
R
E
A
Site 692
T4685
E
A
A
T
W
T
L
T
C
R
P
K
L
C
P
Site 693
T4742
E
G
A
R
A
I
P
T
Y
Q
E
G
A
C
C
Site 694
Y4743
G
A
R
A
I
P
T
Y
Q
E
G
A
C
C
P
Site 695
T4779
V
S
S
S
L
C
E
T
C
R
C
E
L
P
G
Site 696
Y4812
H
C
P
V
G
F
E
Y
Q
E
Q
S
G
Q
C
Site 697
S4835
C
V
T
N
T
S
K
S
P
A
H
L
F
Y
P
Site 698
Y4841
K
S
P
A
H
L
F
Y
P
G
E
T
W
S
D
Site 699
S4878
K
K
A
C
P
P
L
S
C
S
L
D
E
A
R
Site 700
S4880
A
C
P
P
L
S
C
S
L
D
E
A
R
M
S
Site 701
S4887
S
L
D
E
A
R
M
S
K
D
G
C
C
R
F
Site 702
Y4902
C
P
L
P
P
P
P
Y
Q
N
Q
S
T
C
A
Site 703
Y4911
N
Q
S
T
C
A
V
Y
H
R
S
L
I
I
Q
Site 704
S4925
Q
Q
Q
G
C
S
S
S
E
P
V
R
L
A
Y
Site 705
Y4932
S
E
P
V
R
L
A
Y
C
R
G
N
C
G
D
Site 706
S4940
C
R
G
N
C
G
D
S
S
S
M
Y
S
L
E
Site 707
S4941
R
G
N
C
G
D
S
S
S
M
Y
S
L
E
G
Site 708
S4942
G
N
C
G
D
S
S
S
M
Y
S
L
E
G
N
Site 709
Y4944
C
G
D
S
S
S
M
Y
S
L
E
G
N
T
V
Site 710
S4945
G
D
S
S
S
M
Y
S
L
E
G
N
T
V
E
Site 711
S4964
C
C
Q
E
L
R
T
S
L
R
N
V
T
L
H
Site 712
T4969
R
T
S
L
R
N
V
T
L
H
C
T
D
G
S
Site 713
T4973
R
N
V
T
L
H
C
T
D
G
S
S
R
A
F
Site 714
S4976
T
L
H
C
T
D
G
S
S
R
A
F
S
Y
T
Site 715
S4977
L
H
C
T
D
G
S
S
R
A
F
S
Y
T
E
Site 716
S4981
D
G
S
S
R
A
F
S
Y
T
E
V
E
E
C
Site 717
Y4982
G
S
S
R
A
F
S
Y
T
E
V
E
E
C
G
Site 718
T4983
S
S
R
A
F
S
Y
T
E
V
E
E
C
G
C
Site 719
T5001
R
C
P
A
P
G
D
T
Q
H
S
E
E
A
E
Site 720
S5004
A
P
G
D
T
Q
H
S
E
E
A
E
P
E
P
Site 721
S5012
E
E
A
E
P
E
P
S
Q
E
A
E
S
G
S
Site 722
S5019
S
Q
E
A
E
S
G
S
W
E
R
G
V
P
V
Legend
Confirmed in mammals
Confirmed in related proteins or other species
Predicted by Kinexus P-Site Prediction algorithm
No data/link available
Link available
Products available
2019 Kinexus Bioinformatics Corporation