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Updated November 2019
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Protein Info
Short Name:
GON4
Full Name:
GON-4-like protein
Alias:
FLJ20203; GON-4; GON4L; Gon-4-like
Type:
Mass (Da):
248620
Number AA:
2241
UniProt ID:
Q3T8J9
International Prot ID:
IPI00375803
Sequence:
Retrieve full protein sequence
GO Terms
External Links
Internal Links
Cellular Component:
GO:0005737
GO:0005634
Uniprot
OncoNet
Molecular Function:
GO:0003677
PhosphoSite+
KinaseNET
Biological Process:
GO:0006355
Phosida
TranscriptoNet
STRING
Kinexus Products
Info Box
A variety of parameters were considered in the selection of putative P-sites. Confirmed P-Sites have lower Hydrophobicity Scores. The P-site Similarity Score is lower the more that it resembles typical confirmed corresponding P-Ser, P-Thr or P-Tyr sites. The Maximum KInase Score provides the calculated score for highest match of 500 human protein kinases for the amino acid sequence surrounding the target P-site as determined with Kinase Substrate Predictor V2. The Sum KInase Score provides the additive sum of the positive individual Kinase Substrate Predictor V2 scores from 500 human protein kinases. The Conservation Score is the average of the percent similarity of the human P-site with the equivalent P-site in 20 other diverse species. Click the coloured buttons below to retrieve this and other information in the Info Box, or click the orange buttons for relevant links.
Phosphosites
-7
-6
-5
-4
-3
-2
-1
0
1
2
3
4
5
6
7
Expt. conf.
Effect
Kinase
PPase
Kinexus Products
Ref.
Evol.
Kinase Pred.
P-site Match
Site 1
T9
L
P
C
K
K
R
R
T
T
V
T
E
S
L
Q
Site 2
T10
P
C
K
K
R
R
T
T
V
T
E
S
L
Q
H
Site 3
T12
K
K
R
R
T
T
V
T
E
S
L
Q
H
K
G
Site 4
S30
E
N
N
V
D
L
E
S
A
V
K
P
E
S
D
Site 5
S36
E
S
A
V
K
P
E
S
D
Q
V
K
D
L
S
Site 6
S43
S
D
Q
V
K
D
L
S
S
V
S
L
S
W
D
Site 7
S44
D
Q
V
K
D
L
S
S
V
S
L
S
W
D
P
Site 8
S46
V
K
D
L
S
S
V
S
L
S
W
D
P
S
H
Site 9
S48
D
L
S
S
V
S
L
S
W
D
P
S
H
G
R
Site 10
S52
V
S
L
S
W
D
P
S
H
G
R
V
A
G
F
Site 11
S63
V
A
G
F
E
V
Q
S
L
Q
D
A
G
N
Q
Site 12
T76
N
Q
L
G
M
E
D
T
S
L
S
S
G
M
L
Site 13
S77
Q
L
G
M
E
D
T
S
L
S
S
G
M
L
T
Site 14
S79
G
M
E
D
T
S
L
S
S
G
M
L
T
Q
N
Site 15
T84
S
L
S
S
G
M
L
T
Q
N
T
N
V
P
I
Site 16
T87
S
G
M
L
T
Q
N
T
N
V
P
I
L
E
G
Site 17
S107
S
Q
G
I
T
L
P
S
L
E
S
F
H
P
L
Site 18
T126
G
K
G
K
L
H
A
T
G
S
K
R
G
K
K
Site 19
S128
G
K
L
H
A
T
G
S
K
R
G
K
K
M
T
Site 20
T135
S
K
R
G
K
K
M
T
L
R
P
G
P
V
T
Site 21
T142
T
L
R
P
G
P
V
T
Q
E
D
R
C
D
H
Site 22
T151
E
D
R
C
D
H
L
T
L
K
E
P
F
S
G
Site 23
S157
L
T
L
K
E
P
F
S
G
E
P
S
E
E
V
Site 24
S161
E
P
F
S
G
E
P
S
E
E
V
K
E
E
G
Site 25
S175
G
G
K
P
Q
M
N
S
E
G
E
I
P
S
L
Site 26
S181
N
S
E
G
E
I
P
S
L
P
S
G
S
Q
S
Site 27
S184
G
E
I
P
S
L
P
S
G
S
Q
S
A
K
P
Site 28
S186
I
P
S
L
P
S
G
S
Q
S
A
K
P
V
S
Site 29
S188
S
L
P
S
G
S
Q
S
A
K
P
V
S
Q
P
Site 30
S193
S
Q
S
A
K
P
V
S
Q
P
R
K
S
T
Q
Site 31
S198
P
V
S
Q
P
R
K
S
T
Q
P
D
V
C
A
Site 32
T199
V
S
Q
P
R
K
S
T
Q
P
D
V
C
A
S
Site 33
S206
T
Q
P
D
V
C
A
S
P
Q
E
K
P
L
R
Site 34
T250
R
R
K
K
K
K
G
T
K
R
K
R
D
G
R
Site 35
T262
D
G
R
G
Q
E
G
T
L
A
Y
D
L
K
L
Site 36
T276
L
D
D
M
L
D
R
T
L
E
D
G
A
K
Q
Site 37
T326
P
M
F
E
P
K
M
T
R
S
K
L
K
E
V
Site 38
T342
E
K
G
V
V
I
P
T
W
N
I
S
P
I
K
Site 39
S346
V
I
P
T
W
N
I
S
P
I
K
K
A
N
E
Site 40
S369
I
H
L
E
E
D
D
S
S
D
E
E
Y
Q
P
Site 41
S370
H
L
E
E
D
D
S
S
D
E
E
Y
Q
P
D
Site 42
Y374
D
D
S
S
D
E
E
Y
Q
P
D
D
E
E
E
Site 43
T384
D
D
E
E
E
D
E
T
A
E
E
S
L
L
E
Site 44
S388
E
D
E
T
A
E
E
S
L
L
E
S
D
V
E
Site 45
S392
A
E
E
S
L
L
E
S
D
V
E
S
T
A
S
Site 46
S396
L
L
E
S
D
V
E
S
T
A
S
S
P
R
G
Site 47
S399
S
D
V
E
S
T
A
S
S
P
R
G
A
K
K
Site 48
S400
D
V
E
S
T
A
S
S
P
R
G
A
K
K
S
Site 49
S407
S
P
R
G
A
K
K
S
R
L
R
Q
S
S
E
Site 50
S412
K
K
S
R
L
R
Q
S
S
E
M
T
E
T
D
Site 51
S413
K
S
R
L
R
Q
S
S
E
M
T
E
T
D
E
Site 52
T416
L
R
Q
S
S
E
M
T
E
T
D
E
E
S
G
Site 53
T418
Q
S
S
E
M
T
E
T
D
E
E
S
G
I
L
Site 54
S422
M
T
E
T
D
E
E
S
G
I
L
S
E
A
E
Site 55
S426
D
E
E
S
G
I
L
S
E
A
E
K
V
T
T
Site 56
T433
S
E
A
E
K
V
T
T
P
A
I
R
H
I
S
Site 57
S460
K
P
K
Q
T
R
D
S
T
F
M
E
K
L
H
Site 58
T461
P
K
Q
T
R
D
S
T
F
M
E
K
L
H
A
Site 59
S475
A
V
D
E
E
L
A
S
S
P
V
C
M
D
S
Site 60
S482
S
S
P
V
C
M
D
S
F
Q
P
M
D
D
S
Site 61
S489
S
F
Q
P
M
D
D
S
L
I
A
F
R
T
R
Site 62
T495
D
S
L
I
A
F
R
T
R
S
K
M
P
L
K
Site 63
S497
L
I
A
F
R
T
R
S
K
M
P
L
K
D
V
Site 64
T519
E
L
Q
A
P
D
I
T
P
D
M
Y
D
P
N
Site 65
Y523
P
D
I
T
P
D
M
Y
D
P
N
T
A
D
D
Site 66
Y559
D
D
D
D
D
P
E
Y
N
F
L
E
D
L
D
Site 67
T570
E
D
L
D
E
P
D
T
E
D
F
R
T
D
R
Site 68
T575
P
D
T
E
D
F
R
T
D
R
A
V
R
I
T
Site 69
T582
T
D
R
A
V
R
I
T
K
K
E
V
N
E
L
Site 70
T596
L
M
E
E
L
F
E
T
F
Q
D
E
M
G
F
Site 71
T626
E
P
R
P
N
F
N
T
P
Q
A
L
R
F
E
Site 72
T646
L
L
N
E
Q
H
R
T
V
K
E
L
F
E
Q
Site 73
S659
E
Q
L
K
M
K
K
S
S
A
K
Q
L
Q
E
Site 74
T722
L
N
P
E
A
T
T
T
R
I
F
L
K
E
L
Site 75
S736
L
G
T
F
A
Q
S
S
I
A
L
H
H
Q
Y
Site 76
Y743
S
I
A
L
H
H
Q
Y
N
P
K
F
Q
T
L
Site 77
T768
Q
L
I
E
D
F
S
T
H
V
S
I
D
C
S
Site 78
S771
E
D
F
S
T
H
V
S
I
D
C
S
P
H
K
Site 79
S775
T
H
V
S
I
D
C
S
P
H
K
T
V
K
K
Site 80
T779
I
D
C
S
P
H
K
T
V
K
K
T
A
N
E
Site 81
T783
P
H
K
T
V
K
K
T
A
N
E
F
P
C
L
Site 82
S814
P
E
L
L
P
V
C
S
L
K
A
K
N
P
Q
Site 83
S852
E
F
P
N
P
L
I
S
K
Y
L
L
T
C
K
Site 84
T860
K
Y
L
L
T
C
K
T
A
H
Q
L
T
V
R
Site 85
T865
C
K
T
A
H
Q
L
T
V
R
I
K
N
L
N
Site 86
S923
L
P
F
W
L
K
A
S
L
P
S
I
Q
E
E
Site 87
S952
T
G
T
T
E
I
N
S
D
R
S
L
E
K
D
Site 88
S955
T
E
I
N
S
D
R
S
L
E
K
D
N
L
E
Site 89
S965
K
D
N
L
E
L
G
S
E
S
R
Y
P
L
L
Site 90
S967
N
L
E
L
G
S
E
S
R
Y
P
L
L
L
P
Site 91
Y969
E
L
G
S
E
S
R
Y
P
L
L
L
P
K
G
Site 92
S998
K
A
W
R
Q
K
R
S
S
V
L
K
P
L
L
Site 93
S999
A
W
R
Q
K
R
S
S
V
L
K
P
L
L
I
Site 94
S1009
K
P
L
L
I
Q
P
S
P
S
L
Q
P
S
F
Site 95
S1011
L
L
I
Q
P
S
P
S
L
Q
P
S
F
N
P
Site 96
S1015
P
S
P
S
L
Q
P
S
F
N
P
G
K
T
P
Site 97
T1021
P
S
F
N
P
G
K
T
P
A
R
S
T
H
S
Site 98
T1026
G
K
T
P
A
R
S
T
H
S
E
A
P
P
S
Site 99
S1028
T
P
A
R
S
T
H
S
E
A
P
P
S
K
M
Site 100
S1033
T
H
S
E
A
P
P
S
K
M
V
L
R
I
P
Site 101
T1047
P
H
P
I
Q
P
A
T
V
L
Q
T
V
P
G
Site 102
S1061
G
V
P
P
L
G
V
S
G
G
E
S
F
E
S
Site 103
S1068
S
G
G
E
S
F
E
S
P
A
A
L
P
A
V
Site 104
S1082
V
P
P
E
A
R
T
S
F
P
L
S
E
S
Q
Site 105
S1086
A
R
T
S
F
P
L
S
E
S
Q
T
L
L
S
Site 106
S1088
T
S
F
P
L
S
E
S
Q
T
L
L
S
S
A
Site 107
S1093
S
E
S
Q
T
L
L
S
S
A
P
V
P
K
V
Site 108
S1094
E
S
Q
T
L
L
S
S
A
P
V
P
K
V
M
Site 109
S1108
M
L
P
S
L
A
P
S
K
F
R
K
P
Y
V
Site 110
Y1114
P
S
K
F
R
K
P
Y
V
R
R
R
P
S
K
Site 111
S1120
P
Y
V
R
R
R
P
S
K
R
R
G
V
K
A
Site 112
S1128
K
R
R
G
V
K
A
S
P
C
M
K
P
A
P
Site 113
S1219
S
V
N
L
P
I
P
S
T
P
E
D
K
A
H
Site 114
T1220
V
N
L
P
I
P
S
T
P
E
D
K
A
H
V
Site 115
S1255
K
L
E
P
Q
E
L
S
P
L
S
A
T
V
F
Site 116
S1258
P
Q
E
L
S
P
L
S
A
T
V
F
P
K
V
Site 117
T1260
E
L
S
P
L
S
A
T
V
F
P
K
V
E
H
Site 118
S1268
V
F
P
K
V
E
H
S
P
G
P
P
L
A
D
Site 119
S1283
A
E
C
Q
E
G
L
S
E
N
S
A
C
R
W
Site 120
T1291
E
N
S
A
C
R
W
T
V
V
K
T
E
E
G
Site 121
S1312
L
P
Q
G
I
Q
E
S
L
N
N
P
T
P
G
Site 122
T1317
Q
E
S
L
N
N
P
T
P
G
D
L
E
E
I
Site 123
S1339
A
R
E
E
I
S
G
S
P
E
R
D
I
C
D
Site 124
T1358
E
H
A
V
E
L
D
T
G
A
P
S
E
E
L
Site 125
S1362
E
L
D
T
G
A
P
S
E
E
L
S
S
A
G
Site 126
S1366
G
A
P
S
E
E
L
S
S
A
G
E
V
T
K
Site 127
S1367
A
P
S
E
E
L
S
S
A
G
E
V
T
K
Q
Site 128
T1375
A
G
E
V
T
K
Q
T
V
L
Q
K
E
E
E
Site 129
S1384
L
Q
K
E
E
E
R
S
Q
P
T
K
T
P
S
Site 130
T1387
E
E
E
R
S
Q
P
T
K
T
P
S
S
S
Q
Site 131
T1389
E
R
S
Q
P
T
K
T
P
S
S
S
Q
E
P
Site 132
S1391
S
Q
P
T
K
T
P
S
S
S
Q
E
P
P
D
Site 133
S1392
Q
P
T
K
T
P
S
S
S
Q
E
P
P
D
E
Site 134
S1393
P
T
K
T
P
S
S
S
Q
E
P
P
D
E
G
Site 135
T1404
P
D
E
G
T
S
G
T
D
V
N
K
G
S
S
Site 136
S1411
T
D
V
N
K
G
S
S
K
N
A
L
S
S
M
Site 137
S1416
G
S
S
K
N
A
L
S
S
M
D
P
E
V
R
Site 138
S1417
S
S
K
N
A
L
S
S
M
D
P
E
V
R
L
Site 139
S1425
M
D
P
E
V
R
L
S
S
P
P
G
K
P
E
Site 140
S1426
D
P
E
V
R
L
S
S
P
P
G
K
P
E
D
Site 141
S1434
P
P
G
K
P
E
D
S
S
S
V
D
G
Q
S
Site 142
S1435
P
G
K
P
E
D
S
S
S
V
D
G
Q
S
V
Site 143
S1436
G
K
P
E
D
S
S
S
V
D
G
Q
S
V
G
Site 144
S1441
S
S
S
V
D
G
Q
S
V
G
T
P
V
G
P
Site 145
T1444
V
D
G
Q
S
V
G
T
P
V
G
P
E
T
G
Site 146
T1469
E
E
D
F
D
D
L
T
Q
D
E
E
D
E
M
Site 147
S1477
Q
D
E
E
D
E
M
S
S
A
S
E
E
S
V
Site 148
S1478
D
E
E
D
E
M
S
S
A
S
E
E
S
V
L
Site 149
S1480
E
D
E
M
S
S
A
S
E
E
S
V
L
S
V
Site 150
S1483
M
S
S
A
S
E
E
S
V
L
S
V
P
E
L
Site 151
S1486
A
S
E
E
S
V
L
S
V
P
E
L
Q
E
T
Site 152
T1493
S
V
P
E
L
Q
E
T
M
E
K
L
T
W
L
Site 153
S1502
E
K
L
T
W
L
A
S
E
R
R
M
S
Q
E
Site 154
S1507
L
A
S
E
R
R
M
S
Q
E
G
E
S
E
E
Site 155
S1512
R
M
S
Q
E
G
E
S
E
E
E
N
S
Q
E
Site 156
S1517
G
E
S
E
E
E
N
S
Q
E
E
N
S
E
P
Site 157
S1537
E
E
A
E
G
M
E
S
L
Q
K
E
D
E
M
Site 158
T1545
L
Q
K
E
D
E
M
T
D
E
A
V
G
D
S
Site 159
T1558
D
S
A
E
K
P
P
T
F
A
S
P
E
T
A
Site 160
S1561
E
K
P
P
T
F
A
S
P
E
T
A
P
E
V
Site 161
S1571
T
A
P
E
V
E
T
S
R
T
P
P
G
E
S
Site 162
T1573
P
E
V
E
T
S
R
T
P
P
G
E
S
I
K
Site 163
S1597
R
A
R
N
K
R
G
S
R
A
R
A
S
K
D
Site 164
S1602
R
G
S
R
A
R
A
S
K
D
T
S
K
L
L
Site 165
T1605
R
A
R
A
S
K
D
T
S
K
L
L
L
L
Y
Site 166
S1606
A
R
A
S
K
D
T
S
K
L
L
L
L
Y
D
Site 167
Y1612
T
S
K
L
L
L
L
Y
D
E
D
I
L
E
R
Site 168
Y1649
L
Q
H
I
P
G
K
Y
E
D
F
L
Q
V
I
Site 169
Y1657
E
D
F
L
Q
V
I
Y
E
F
E
S
S
T
Q
Site 170
S1662
V
I
Y
E
F
E
S
S
T
Q
R
R
T
A
V
Site 171
T1663
I
Y
E
F
E
S
S
T
Q
R
R
T
A
V
D
Site 172
T1667
E
S
S
T
Q
R
R
T
A
V
D
L
Y
K
S
Site 173
S1713
E
Q
Q
A
F
E
K
S
R
K
F
L
R
Q
L
Site 174
T1793
D
F
E
E
I
N
W
T
E
E
K
E
Y
E
F
Site 175
Y1798
N
W
T
E
E
K
E
Y
E
F
D
G
F
E
E
Site 176
T1841
V
Q
N
H
D
K
E
T
E
W
P
D
G
A
K
Site 177
S1861
C
H
E
G
G
P
D
S
K
L
K
K
S
K
R
Site 178
S1866
P
D
S
K
L
K
K
S
K
R
R
S
C
S
H
Site 179
S1870
L
K
K
S
K
R
R
S
C
S
H
C
S
S
K
Site 180
S1872
K
S
K
R
R
S
C
S
H
C
S
S
K
V
C
Site 181
S1875
R
R
S
C
S
H
C
S
S
K
V
C
D
S
K
Site 182
S1876
R
S
C
S
H
C
S
S
K
V
C
D
S
K
S
Site 183
S1881
C
S
S
K
V
C
D
S
K
S
Y
K
S
K
E
Site 184
S1883
S
K
V
C
D
S
K
S
Y
K
S
K
E
P
H
Site 185
Y1884
K
V
C
D
S
K
S
Y
K
S
K
E
P
H
E
Site 186
S1886
C
D
S
K
S
Y
K
S
K
E
P
H
E
L
V
Site 187
S1895
E
P
H
E
L
V
G
S
S
P
H
R
E
A
S
Site 188
S1896
P
H
E
L
V
G
S
S
P
H
R
E
A
S
P
Site 189
S1902
S
S
P
H
R
E
A
S
P
M
P
G
A
K
E
Site 190
S1927
E
A
P
E
E
R
E
S
T
E
A
T
Q
S
R
Site 191
T1928
A
P
E
E
R
E
S
T
E
A
T
Q
S
R
T
Site 192
T1931
E
R
E
S
T
E
A
T
Q
S
R
T
V
R
T
Site 193
S1933
E
S
T
E
A
T
Q
S
R
T
V
R
T
T
R
Site 194
T1935
T
E
A
T
Q
S
R
T
V
R
T
T
R
K
G
Site 195
T1939
Q
S
R
T
V
R
T
T
R
K
G
E
M
P
V
Site 196
S1947
R
K
G
E
M
P
V
S
A
G
L
A
V
G
S
Site 197
S1954
S
A
G
L
A
V
G
S
T
L
P
S
P
R
E
Site 198
T1965
S
P
R
E
V
T
V
T
E
R
L
L
L
D
G
Site 199
S1977
L
D
G
P
P
P
H
S
P
E
T
P
Q
F
P
Site 200
T1980
P
P
P
H
S
P
E
T
P
Q
F
P
P
T
T
Site 201
T1986
E
T
P
Q
F
P
P
T
T
G
A
V
L
Y
T
Site 202
Y1992
P
T
T
G
A
V
L
Y
T
V
K
R
N
Q
V
Site 203
T1993
T
T
G
A
V
L
Y
T
V
K
R
N
Q
V
G
Site 204
S2005
Q
V
G
P
E
V
R
S
C
P
K
A
S
P
R
Site 205
S2010
V
R
S
C
P
K
A
S
P
R
L
Q
K
E
R
Site 206
S2036
L
M
L
V
W
D
A
S
E
T
E
K
L
P
G
Site 207
T2044
E
T
E
K
L
P
G
T
V
E
P
P
A
S
F
Site 208
S2053
E
P
P
A
S
F
L
S
P
V
S
S
K
T
R
Site 209
S2056
A
S
F
L
S
P
V
S
S
K
T
R
D
A
G
Site 210
S2057
S
F
L
S
P
V
S
S
K
T
R
D
A
G
R
Site 211
S2068
D
A
G
R
R
H
V
S
G
K
P
D
T
Q
E
Site 212
T2073
H
V
S
G
K
P
D
T
Q
E
R
W
L
P
S
Site 213
S2080
T
Q
E
R
W
L
P
S
S
R
A
R
V
K
T
Site 214
S2081
Q
E
R
W
L
P
S
S
R
A
R
V
K
T
R
Site 215
T2087
S
S
R
A
R
V
K
T
R
D
R
T
C
P
V
Site 216
T2091
R
V
K
T
R
D
R
T
C
P
V
H
E
S
P
Site 217
S2097
R
T
C
P
V
H
E
S
P
S
G
I
D
T
S
Site 218
S2099
C
P
V
H
E
S
P
S
G
I
D
T
S
E
T
Site 219
T2103
E
S
P
S
G
I
D
T
S
E
T
S
P
K
A
Site 220
S2104
S
P
S
G
I
D
T
S
E
T
S
P
K
A
P
Site 221
T2106
S
G
I
D
T
S
E
T
S
P
K
A
P
R
G
Site 222
S2107
G
I
D
T
S
E
T
S
P
K
A
P
R
G
G
Site 223
S2119
R
G
G
L
A
K
D
S
G
T
Q
A
K
G
P
Site 224
T2121
G
L
A
K
D
S
G
T
Q
A
K
G
P
E
G
Site 225
S2144
A
T
V
C
A
N
N
S
K
V
S
S
T
G
E
Site 226
S2147
C
A
N
N
S
K
V
S
S
T
G
E
K
V
V
Site 227
S2148
A
N
N
S
K
V
S
S
T
G
E
K
V
V
L
Site 228
T2157
G
E
K
V
V
L
W
T
R
E
A
D
R
V
I
Site 229
T2166
E
A
D
R
V
I
L
T
M
C
Q
E
Q
G
A
Site 230
T2177
E
Q
G
A
Q
P
Q
T
F
N
I
I
S
Q
Q
Site 231
S2182
P
Q
T
F
N
I
I
S
Q
Q
L
G
N
K
T
Site 232
T2189
S
Q
Q
L
G
N
K
T
P
A
E
V
S
H
R
Site 233
S2194
N
K
T
P
A
E
V
S
H
R
F
R
E
L
M
Site 234
T2206
E
L
M
Q
L
F
H
T
A
C
E
A
S
S
E
Site 235
T2219
S
E
D
E
D
D
A
T
S
T
S
N
A
D
Q
Site 236
S2222
E
D
D
A
T
S
T
S
N
A
D
Q
L
S
D
Site 237
S2228
T
S
N
A
D
Q
L
S
D
H
G
D
L
L
S
Site 238
S2235
S
D
H
G
D
L
L
S
E
E
E
L
D
E
_
Legend
Confirmed in mammals
Confirmed in related proteins or other species
Predicted by Kinexus P-Site Prediction algorithm
No data/link available
Link available
Products available
2019 Kinexus Bioinformatics Corporation