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Search by protein name, UniProt number, IPI number, or 15 AA P-site sequence.
Updated November 2019
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Protein Info
Short Name:
DDX4
Full Name:
Probable ATP-dependent RNA helicase DDX4
Alias:
ATP-dependent RNA helicase DDX4; DEAD box 4; EC 3.6.1.-; Vasa
Type:
Ribonucleoprotein complex, Cytoplasm, Nucleus protein
Mass (Da):
79308
Number AA:
724
UniProt ID:
Q9NQI0
International Prot ID:
IPI00071483
Sequence:
Retrieve full protein sequence
GO Terms
External Links
Internal Links
Cellular Component:
GO:0005737
Uniprot
OncoNet
Molecular Function:
GO:0005524
GO:0008026
GO:0003676
PhosphoSite+
KinaseNET
Biological Process:
GO:0007275
Phosida
TranscriptoNet
STRING
Kinexus Products
Info Box
A variety of parameters were considered in the selection of putative P-sites. Confirmed P-Sites have lower Hydrophobicity Scores. The P-site Similarity Score is lower the more that it resembles typical confirmed corresponding P-Ser, P-Thr or P-Tyr sites. The Maximum KInase Score provides the calculated score for highest match of 500 human protein kinases for the amino acid sequence surrounding the target P-site as determined with Kinase Substrate Predictor V2. The Sum KInase Score provides the additive sum of the positive individual Kinase Substrate Predictor V2 scores from 500 human protein kinases. The Conservation Score is the average of the percent similarity of the human P-site with the equivalent P-site in 20 other diverse species. Click the coloured buttons below to retrieve this and other information in the Info Box, or click the orange buttons for relevant links.
Phosphosites
-7
-6
-5
-4
-3
-2
-1
0
1
2
3
4
5
6
7
Expt. conf.
Effect
Kinase
PPase
Kinexus Products
Ref.
Evol.
Kinase Pred.
P-site Match
Site 1
S16
E
I
N
P
H
M
S
S
Y
V
P
I
F
E
K
Site 2
Y17
I
N
P
H
M
S
S
Y
V
P
I
F
E
K
D
Site 3
Y26
P
I
F
E
K
D
R
Y
S
G
E
N
G
D
N
Site 4
S27
I
F
E
K
D
R
Y
S
G
E
N
G
D
N
F
Site 5
T37
N
G
D
N
F
N
R
T
P
A
S
S
S
E
M
Site 6
S40
N
F
N
R
T
P
A
S
S
S
E
M
D
D
G
Site 7
S41
F
N
R
T
P
A
S
S
S
E
M
D
D
G
P
Site 8
S42
N
R
T
P
A
S
S
S
E
M
D
D
G
P
S
Site 9
S49
S
E
M
D
D
G
P
S
R
R
D
H
F
M
K
Site 10
S57
R
R
D
H
F
M
K
S
G
F
A
S
G
R
N
Site 11
S61
F
M
K
S
G
F
A
S
G
R
N
F
G
N
R
Site 12
T79
E
C
N
K
R
D
N
T
S
T
M
G
G
F
G
Site 13
T81
N
K
R
D
N
T
S
T
M
G
G
F
G
V
G
Site 14
S90
G
G
F
G
V
G
K
S
F
G
N
R
G
F
S
Site 15
S97
S
F
G
N
R
G
F
S
N
S
R
F
E
D
G
Site 16
S99
G
N
R
G
F
S
N
S
R
F
E
D
G
D
S
Site 17
S106
S
R
F
E
D
G
D
S
S
G
F
W
R
E
S
Site 18
S107
R
F
E
D
G
D
S
S
G
F
W
R
E
S
S
Site 19
S113
S
S
G
F
W
R
E
S
S
N
D
C
E
D
N
Site 20
S114
S
G
F
W
R
E
S
S
N
D
C
E
D
N
P
Site 21
T122
N
D
C
E
D
N
P
T
R
N
R
G
F
S
K
Site 22
S128
P
T
R
N
R
G
F
S
K
R
G
G
Y
R
D
Site 23
Y133
G
F
S
K
R
G
G
Y
R
D
G
N
N
S
E
Site 24
S139
G
Y
R
D
G
N
N
S
E
A
S
G
P
Y
R
Site 25
S142
D
G
N
N
S
E
A
S
G
P
Y
R
R
G
G
Site 26
S152
Y
R
R
G
G
R
G
S
F
R
G
C
R
G
G
Site 27
S164
R
G
G
F
G
L
G
S
P
N
N
D
L
D
P
Site 28
S184
R
T
G
G
L
F
G
S
R
R
P
V
L
S
G
Site 29
S190
G
S
R
R
P
V
L
S
G
T
G
N
G
D
T
Site 30
T197
S
G
T
G
N
G
D
T
S
Q
S
R
S
G
S
Site 31
S198
G
T
G
N
G
D
T
S
Q
S
R
S
G
S
G
Site 32
S200
G
N
G
D
T
S
Q
S
R
S
G
S
G
S
E
Site 33
S202
G
D
T
S
Q
S
R
S
G
S
G
S
E
R
G
Site 34
S204
T
S
Q
S
R
S
G
S
G
S
E
R
G
G
Y
Site 35
S206
Q
S
R
S
G
S
G
S
E
R
G
G
Y
K
G
Site 36
Y211
S
G
S
E
R
G
G
Y
K
G
L
N
E
E
V
Site 37
T220
G
L
N
E
E
V
I
T
G
S
G
K
N
S
W
Site 38
S222
N
E
E
V
I
T
G
S
G
K
N
S
W
K
S
Site 39
S226
I
T
G
S
G
K
N
S
W
K
S
E
A
E
G
Site 40
S229
S
G
K
N
S
W
K
S
E
A
E
G
G
E
S
Site 41
S236
S
E
A
E
G
G
E
S
S
D
T
Q
G
P
K
Site 42
S237
E
A
E
G
G
E
S
S
D
T
Q
G
P
K
V
Site 43
T239
E
G
G
E
S
S
D
T
Q
G
P
K
V
T
Y
Site 44
T245
D
T
Q
G
P
K
V
T
Y
I
P
P
P
P
P
Site 45
Y246
T
Q
G
P
K
V
T
Y
I
P
P
P
P
P
E
Site 46
S257
P
P
P
E
D
E
D
S
I
F
A
H
Y
Q
T
Site 47
Y262
E
D
S
I
F
A
H
Y
Q
T
G
I
N
F
D
Site 48
T312
K
A
G
Y
T
K
L
T
P
V
Q
K
Y
S
I
Site 49
T339
A
Q
T
G
S
G
K
T
A
A
F
L
L
P
I
Site 50
T356
H
M
M
H
D
G
I
T
A
S
R
F
K
E
L
Site 51
S358
M
H
D
G
I
T
A
S
R
F
K
E
L
Q
E
Site 52
Y382
R
E
L
V
N
Q
I
Y
L
E
A
R
K
F
S
Site 53
S389
Y
L
E
A
R
K
F
S
F
G
T
C
V
R
A
Site 54
Y400
C
V
R
A
V
V
I
Y
G
G
T
Q
L
G
H
Site 55
S408
G
G
T
Q
L
G
H
S
I
R
Q
I
V
Q
G
Site 56
T422
G
C
N
I
L
C
A
T
P
G
R
L
M
D
I
Site 57
Y442
I
G
L
K
Q
I
K
Y
L
V
L
D
E
A
D
Site 58
S465
P
E
M
K
K
L
I
S
C
P
G
M
P
S
K
Site 59
T477
P
S
K
E
Q
R
Q
T
L
M
F
S
A
T
F
Site 60
T483
Q
T
L
M
F
S
A
T
F
P
E
E
I
Q
R
Site 61
T518
A
C
R
D
V
Q
Q
T
V
L
Q
V
G
Q
F
Site 62
S526
V
L
Q
V
G
Q
F
S
K
R
E
K
L
V
E
Site 63
T543
R
N
I
G
D
E
R
T
M
V
F
V
E
T
K
Site 64
T549
R
T
M
V
F
V
E
T
K
K
K
A
D
F
I
Site 65
T567
L
C
Q
E
K
I
S
T
T
S
I
H
G
D
R
Site 66
T620
I
N
F
D
L
P
S
T
I
D
E
Y
V
H
R
Site 67
Y624
L
P
S
T
I
D
E
Y
V
H
R
I
G
R
T
Site 68
T637
R
T
G
R
C
G
N
T
G
R
A
I
S
F
F
Site 69
S642
G
N
T
G
R
A
I
S
F
F
D
L
E
S
D
Site 70
S648
I
S
F
F
D
L
E
S
D
N
H
L
A
Q
P
Site 71
Y679
E
E
I
A
F
S
T
Y
I
P
G
F
S
G
S
Site 72
S684
S
T
Y
I
P
G
F
S
G
S
T
R
G
N
V
Site 73
S686
Y
I
P
G
F
S
G
S
T
R
G
N
V
F
A
Site 74
S694
T
R
G
N
V
F
A
S
V
D
T
R
K
G
K
Site 75
S702
V
D
T
R
K
G
K
S
T
L
N
T
A
G
F
Site 76
T703
D
T
R
K
G
K
S
T
L
N
T
A
G
F
S
Site 77
T706
K
G
K
S
T
L
N
T
A
G
F
S
S
S
Q
Site 78
S712
N
T
A
G
F
S
S
S
Q
A
P
N
P
V
D
Legend
Confirmed in mammals
Confirmed in related proteins or other species
Predicted by Kinexus P-Site Prediction algorithm
No data/link available
Link available
Products available
2019 Kinexus Bioinformatics Corporation