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Search by protein name, UniProt number, IPI number, or 15 AA P-site sequence.
Updated November 2019
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Protein Info
Short Name:
UBQLN4
Full Name:
Ubiquilin-4
Alias:
A1U; Ataxin-1 ubiquitin-like-interacting A1U; UBIN; UBQL4
Type:
Targeting protein; Endoplasmic reticulum, nucleus, cytoplasm
Mass (Da):
63853
Number AA:
601
UniProt ID:
Q9NRR5
International Prot ID:
IPI00024502
Sequence:
Retrieve full protein sequence
GO Terms
External Links
Internal Links
Cellular Component:
GO:0005829
GO:0005789
GO:0005634
Uniprot
OncoNet
Molecular Function:
GO:0042802
PhosphoSite+
KinaseNET
Biological Process:
Phosida
TranscriptoNet
STRING
Kinexus Products
Info Box
A variety of parameters were considered in the selection of putative P-sites. Confirmed P-Sites have lower Hydrophobicity Scores. The P-site Similarity Score is lower the more that it resembles typical confirmed corresponding P-Ser, P-Thr or P-Tyr sites. The Maximum KInase Score provides the calculated score for highest match of 500 human protein kinases for the amino acid sequence surrounding the target P-site as determined with Kinase Substrate Predictor V2. The Sum KInase Score provides the additive sum of the positive individual Kinase Substrate Predictor V2 scores from 500 human protein kinases. The Conservation Score is the average of the percent similarity of the human P-site with the equivalent P-site in 20 other diverse species. Click the coloured buttons below to retrieve this and other information in the Info Box, or click the orange buttons for relevant links.
Phosphosites
-7
-6
-5
-4
-3
-2
-1
0
1
2
3
4
5
6
7
Expt. conf.
Effect
Kinase
PPase
Kinexus Products
Ref.
Evol.
Kinase Pred.
P-site Match
Site 1
T9
A
E
P
S
G
A
E
T
R
P
P
I
R
V
T
Site 2
T16
T
R
P
P
I
R
V
T
V
K
T
P
K
D
K
Site 3
T19
P
I
R
V
T
V
K
T
P
K
D
K
E
E
I
Site 4
S33
I
V
I
C
D
R
A
S
V
K
E
F
K
E
E
Site 5
S42
K
E
F
K
E
E
I
S
R
R
F
K
A
Q
Q
Site 6
T66
K
I
L
K
D
G
D
T
L
N
Q
H
G
I
K
Site 7
T84
T
V
H
L
V
I
K
T
P
Q
K
A
Q
D
P
Site 8
S97
D
P
A
A
A
T
A
S
S
P
S
T
P
D
P
Site 9
S98
P
A
A
A
T
A
S
S
P
S
T
P
D
P
A
Site 10
T101
A
T
A
S
S
P
S
T
P
D
P
A
S
A
P
Site 11
S106
P
S
T
P
D
P
A
S
A
P
S
T
T
P
A
Site 12
S109
P
D
P
A
S
A
P
S
T
T
P
A
S
P
A
Site 13
T110
D
P
A
S
A
P
S
T
T
P
A
S
P
A
T
Site 14
T111
P
A
S
A
P
S
T
T
P
A
S
P
A
T
P
Site 15
S114
A
P
S
T
T
P
A
S
P
A
T
P
A
Q
P
Site 16
T117
T
T
P
A
S
P
A
T
P
A
Q
P
S
T
S
Site 17
S122
P
A
T
P
A
Q
P
S
T
S
G
S
A
S
S
Site 18
T123
A
T
P
A
Q
P
S
T
S
G
S
A
S
S
D
Site 19
S124
T
P
A
Q
P
S
T
S
G
S
A
S
S
D
A
Site 20
S126
A
Q
P
S
T
S
G
S
A
S
S
D
A
G
S
Site 21
S128
P
S
T
S
G
S
A
S
S
D
A
G
S
G
S
Site 22
S129
S
T
S
G
S
A
S
S
D
A
G
S
G
S
R
Site 23
S133
S
A
S
S
D
A
G
S
G
S
R
R
S
S
G
Site 24
S135
S
S
D
A
G
S
G
S
R
R
S
S
G
G
G
Site 25
S138
A
G
S
G
S
R
R
S
S
G
G
G
P
S
P
Site 26
S139
G
S
G
S
R
R
S
S
G
G
G
P
S
P
G
Site 27
S144
R
S
S
G
G
G
P
S
P
G
A
G
E
G
S
Site 28
S151
S
P
G
A
G
E
G
S
P
S
A
T
A
S
I
Site 29
S153
G
A
G
E
G
S
P
S
A
T
A
S
I
L
S
Site 30
S191
Q
M
Q
R
Q
L
M
S
N
P
E
M
L
S
Q
Site 31
S197
M
S
N
P
E
M
L
S
Q
I
M
E
N
P
L
Site 32
T248
N
P
E
L
M
R
Q
T
M
E
L
A
R
N
P
Site 33
S270
R
N
Q
D
R
A
L
S
N
L
E
S
I
P
G
Site 34
S274
R
A
L
S
N
L
E
S
I
P
G
G
Y
N
A
Site 35
Y279
L
E
S
I
P
G
G
Y
N
A
L
R
R
M
Y
Site 36
Y286
Y
N
A
L
R
R
M
Y
T
D
I
Q
E
P
M
Site 37
T287
N
A
L
R
R
M
Y
T
D
I
Q
E
P
M
F
Site 38
S295
D
I
Q
E
P
M
F
S
A
A
R
E
Q
F
G
Site 39
S307
Q
F
G
N
N
P
F
S
S
L
A
G
N
S
D
Site 40
S308
F
G
N
N
P
F
S
S
L
A
G
N
S
D
S
Site 41
S313
F
S
S
L
A
G
N
S
D
S
S
S
S
Q
P
Site 42
S315
S
L
A
G
N
S
D
S
S
S
S
Q
P
L
R
Site 43
S316
L
A
G
N
S
D
S
S
S
S
Q
P
L
R
T
Site 44
S317
A
G
N
S
D
S
S
S
S
Q
P
L
R
T
E
Site 45
S318
G
N
S
D
S
S
S
S
Q
P
L
R
T
E
N
Site 46
T323
S
S
S
Q
P
L
R
T
E
N
R
E
P
L
P
Site 47
S334
E
P
L
P
N
P
W
S
P
S
P
P
T
S
Q
Site 48
S336
L
P
N
P
W
S
P
S
P
P
T
S
Q
A
P
Site 49
T339
P
W
S
P
S
P
P
T
S
Q
A
P
G
S
G
Site 50
S340
W
S
P
S
P
P
T
S
Q
A
P
G
S
G
G
Site 51
S345
P
T
S
Q
A
P
G
S
G
G
E
G
T
G
G
Site 52
T350
P
G
S
G
G
E
G
T
G
G
S
G
T
S
Q
Site 53
S353
G
G
E
G
T
G
G
S
G
T
S
Q
V
H
P
Site 54
T355
E
G
T
G
G
S
G
T
S
Q
V
H
P
T
V
Site 55
S356
G
T
G
G
S
G
T
S
Q
V
H
P
T
V
S
Site 56
T361
G
T
S
Q
V
H
P
T
V
S
N
P
F
G
I
Site 57
S363
S
Q
V
H
P
T
V
S
N
P
F
G
I
N
A
Site 58
S391
Q
A
L
L
Q
Q
I
S
E
N
P
Q
L
M
Q
Site 59
Y405
Q
N
V
I
S
A
P
Y
M
R
S
M
M
Q
T
Site 60
S456
Q
Q
M
Q
N
P
E
S
L
S
I
L
T
N
P
Site 61
T481
Q
G
L
Q
T
L
Q
T
E
A
P
G
L
V
P
Site 62
S496
S
L
G
S
F
G
I
S
R
T
P
A
P
S
A
Site 63
T498
G
S
F
G
I
S
R
T
P
A
P
S
A
G
S
Site 64
S502
I
S
R
T
P
A
P
S
A
G
S
N
A
G
S
Site 65
S505
T
P
A
P
S
A
G
S
N
A
G
S
T
P
E
Site 66
S509
S
A
G
S
N
A
G
S
T
P
E
A
P
T
S
Site 67
T510
A
G
S
N
A
G
S
T
P
E
A
P
T
S
S
Site 68
S516
S
T
P
E
A
P
T
S
S
P
A
T
P
A
T
Site 69
S517
T
P
E
A
P
T
S
S
P
A
T
P
A
T
S
Site 70
T520
A
P
T
S
S
P
A
T
P
A
T
S
S
P
T
Site 71
T523
S
S
P
A
T
P
A
T
S
S
P
T
G
A
S
Site 72
S524
S
P
A
T
P
A
T
S
S
P
T
G
A
S
S
Site 73
S525
P
A
T
P
A
T
S
S
P
T
G
A
S
S
A
Site 74
S531
S
S
P
T
G
A
S
S
A
Q
Q
Q
L
M
Q
Site 75
S550
L
L
A
G
S
G
N
S
Q
V
Q
T
P
E
V
Site 76
T554
S
G
N
S
Q
V
Q
T
P
E
V
R
F
Q
Q
Site 77
S568
Q
Q
L
E
Q
L
N
S
M
G
F
I
N
R
E
Legend
Confirmed in mammals
Confirmed in related proteins or other species
Predicted by Kinexus P-Site Prediction algorithm
No data/link available
Link available
Products available
2019 Kinexus Bioinformatics Corporation