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Search by protein name, UniProt number, IPI number, or 15 AA P-site sequence.
Updated November 2019
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Protein Info
Short Name:
DNAH9
Full Name:
Dynein heavy chain 9, axonemal
Alias:
Axonemal beta dynein heavy chain 9; Ciliary dynein heavy chain 9
Type:
Mass (Da):
511887
Number AA:
4486
UniProt ID:
Q9NYC9
International Prot ID:
IPI00302453
Sequence:
Retrieve full protein sequence
GO Terms
External Links
Internal Links
Cellular Component:
GO:0005856
GO:0005858
GO:0005874
Uniprot
OncoNet
Molecular Function:
GO:0000166
GO:0003774
GO:0003777
PhosphoSite+
KinaseNET
Biological Process:
GO:0000003
GO:0000902
GO:0006810
Phosida
TranscriptoNet
STRING
Kinexus Products
Info Box
A variety of parameters were considered in the selection of putative P-sites. Confirmed P-Sites have lower Hydrophobicity Scores. The P-site Similarity Score is lower the more that it resembles typical confirmed corresponding P-Ser, P-Thr or P-Tyr sites. The Maximum KInase Score provides the calculated score for highest match of 500 human protein kinases for the amino acid sequence surrounding the target P-site as determined with Kinase Substrate Predictor V2. The Sum KInase Score provides the additive sum of the positive individual Kinase Substrate Predictor V2 scores from 500 human protein kinases. The Conservation Score is the average of the percent similarity of the human P-site with the equivalent P-site in 20 other diverse species. Click the coloured buttons below to retrieve this and other information in the Info Box, or click the orange buttons for relevant links.
Phosphosites
-7
-6
-5
-4
-3
-2
-1
0
1
2
3
4
5
6
7
Expt. conf.
Effect
Kinase
PPase
Kinexus Products
Ref.
Evol.
Kinase Pred.
P-site Match
Site 1
T30
R
R
L
R
L
L
G
T
Y
V
A
M
S
L
R
Site 2
T106
K
A
L
F
F
L
R
T
G
P
E
P
P
G
P
Site 3
S115
P
E
P
P
G
P
D
S
F
R
G
A
V
V
C
Site 4
S168
D
V
R
R
H
A
H
S
L
Q
C
D
L
S
V
Site 5
S192
L
L
P
L
P
A
G
S
E
K
M
E
F
A
D
Site 6
S200
E
K
M
E
F
A
D
S
K
S
E
T
V
L
D
Site 7
S202
M
E
F
A
D
S
K
S
E
T
V
L
D
S
I
Site 8
T204
F
A
D
S
K
S
E
T
V
L
D
S
I
D
K
Site 9
S208
K
S
E
T
V
L
D
S
I
D
K
S
V
I
Y
Site 10
S225
E
S
A
V
I
K
W
S
Y
Q
V
Q
V
V
L
Site 11
S236
Q
V
V
L
K
R
E
S
S
Q
P
L
L
Q
G
Site 12
S237
V
V
L
K
R
E
S
S
Q
P
L
L
Q
G
E
Site 13
T247
L
L
Q
G
E
N
P
T
P
K
V
E
L
E
F
Site 14
S257
V
E
L
E
F
W
K
S
R
Y
E
D
L
K
Y
Site 15
Y259
L
E
F
W
K
S
R
Y
E
D
L
K
Y
I
Y
Site 16
Y264
S
R
Y
E
D
L
K
Y
I
Y
N
Q
L
R
T
Site 17
T273
Y
N
Q
L
R
T
I
T
V
R
G
M
A
K
L
Site 18
S286
K
L
L
D
K
L
Q
S
S
Y
F
P
A
F
K
Site 19
Y288
L
D
K
L
Q
S
S
Y
F
P
A
F
K
A
M
Site 20
S351
L
I
W
A
T
C
K
S
Y
R
S
P
G
R
L
Site 21
Y352
I
W
A
T
C
K
S
Y
R
S
P
G
R
L
T
Site 22
S354
A
T
C
K
S
Y
R
S
P
G
R
L
T
V
L
Site 23
S374
N
L
L
I
Q
Q
A
S
N
Y
L
S
P
E
D
Site 24
Y376
L
I
Q
Q
A
S
N
Y
L
S
P
E
D
L
L
Site 25
S378
Q
Q
A
S
N
Y
L
S
P
E
D
L
L
R
S
Site 26
S385
S
P
E
D
L
L
R
S
E
V
E
E
S
Q
R
Site 27
S390
L
R
S
E
V
E
E
S
Q
R
K
L
Q
V
V
Site 28
S398
Q
R
K
L
Q
V
V
S
D
T
L
S
F
F
K
Site 29
T400
K
L
Q
V
V
S
D
T
L
S
F
F
K
Q
E
Site 30
S402
Q
V
V
S
D
T
L
S
F
F
K
Q
E
F
Q
Site 31
T417
D
R
R
E
N
L
H
T
Y
F
K
E
N
Q
E
Site 32
Y418
R
R
E
N
L
H
T
Y
F
K
E
N
Q
E
V
Site 33
S433
K
E
W
D
F
Q
S
S
L
V
F
V
R
L
D
Site 34
S476
G
V
R
G
N
A
L
S
Q
Q
V
Q
Q
M
H
Site 35
Y490
H
E
E
F
Q
E
M
Y
R
L
L
S
G
S
S
Site 36
S494
Q
E
M
Y
R
L
L
S
G
S
S
S
D
C
L
Site 37
S498
R
L
L
S
G
S
S
S
D
C
L
Y
L
Q
S
Site 38
Y502
G
S
S
S
D
C
L
Y
L
Q
S
T
D
F
E
Site 39
S513
T
D
F
E
N
D
V
S
E
F
N
Q
K
V
E
Site 40
T528
D
L
D
R
R
L
G
T
I
F
I
Q
A
F
D
Site 41
T563
R
P
L
V
A
R
D
T
S
D
K
Y
L
V
L
Site 42
Y567
A
R
D
T
S
D
K
Y
L
V
L
I
Q
M
F
Site 43
Y585
L
D
A
V
R
M
I
Y
S
Q
H
V
Q
E
E
Site 44
S586
D
A
V
R
M
I
Y
S
Q
H
V
Q
E
E
A
Site 45
S598
E
E
A
E
L
G
F
S
P
V
H
K
N
M
P
Site 46
T606
P
V
H
K
N
M
P
T
V
A
G
G
L
R
W
Site 47
S626
Q
R
I
Q
G
P
F
S
N
F
G
R
I
T
H
Site 48
Y648
G
K
R
M
Q
Q
K
Y
E
D
M
L
S
L
L
Site 49
S653
Q
K
Y
E
D
M
L
S
L
L
E
K
Y
E
T
Site 50
Y658
M
L
S
L
L
E
K
Y
E
T
R
L
Y
E
D
Site 51
T660
S
L
L
E
K
Y
E
T
R
L
Y
E
D
W
C
Site 52
Y663
E
K
Y
E
T
R
L
Y
E
D
W
C
R
T
V
Site 53
T669
L
Y
E
D
W
C
R
T
V
S
E
K
S
Q
Y
Site 54
S671
E
D
W
C
R
T
V
S
E
K
S
Q
Y
N
L
Site 55
S674
C
R
T
V
S
E
K
S
Q
Y
N
L
S
Q
P
Site 56
Y676
T
V
S
E
K
S
Q
Y
N
L
S
Q
P
L
L
Site 57
S679
E
K
S
Q
Y
N
L
S
Q
P
L
L
K
R
D
Site 58
T689
L
L
K
R
D
P
E
T
K
E
I
T
I
N
F
Site 59
S702
N
F
N
P
Q
L
I
S
V
L
K
E
M
S
Y
Site 60
S708
I
S
V
L
K
E
M
S
Y
L
E
P
R
E
M
Site 61
Y709
S
V
L
K
E
M
S
Y
L
E
P
R
E
M
K
Site 62
S727
E
T
A
A
A
M
F
S
S
R
D
F
Y
R
Q
Site 63
Y732
M
F
S
S
R
D
F
Y
R
Q
L
V
A
N
L
Site 64
Y746
L
E
L
M
A
N
W
Y
N
K
V
M
K
T
L
Site 65
T778
R
L
R
A
A
E
E
T
L
N
W
K
T
E
G
Site 66
Y789
K
T
E
G
I
C
D
Y
V
T
E
I
T
S
S
Site 67
T791
E
G
I
C
D
Y
V
T
E
I
T
S
S
I
H
Site 68
S796
Y
V
T
E
I
T
S
S
I
H
D
L
E
Q
R
Site 69
T823
N
I
M
K
T
W
V
T
P
I
F
K
T
K
D
Site 70
S835
T
K
D
G
K
R
E
S
L
L
S
L
D
D
R
Site 71
S838
G
K
R
E
S
L
L
S
L
D
D
R
H
D
R
Site 72
Y849
R
H
D
R
M
E
K
Y
Y
N
L
I
K
E
S
Site 73
Y850
H
D
R
M
E
K
Y
Y
N
L
I
K
E
S
G
Site 74
S872
Q
E
N
L
G
L
F
S
A
D
P
T
S
N
I
Site 75
T876
G
L
F
S
A
D
P
T
S
N
I
W
K
T
Y
Site 76
Y883
T
S
N
I
W
K
T
Y
V
N
S
I
D
N
L
Site 77
S936
P
E
L
V
F
Y
P
S
L
E
S
G
V
K
G
Site 78
S967
P
S
L
V
P
R
L
S
P
Q
N
G
S
P
H
Site 79
S972
R
L
S
P
Q
N
G
S
P
H
Y
Q
V
D
L
Site 80
Y975
P
Q
N
G
S
P
H
Y
Q
V
D
L
D
G
I
Site 81
T991
D
L
A
N
M
R
R
T
L
M
E
R
V
Q
R
Site 82
Y1015
S
T
F
S
Q
Y
S
Y
L
Y
V
E
D
R
K
Site 83
Y1017
F
S
Q
Y
S
Y
L
Y
V
E
D
R
K
E
V
Site 84
T1036
L
L
Y
G
H
I
L
T
P
E
E
I
E
D
H
Site 85
S1056
P
E
N
P
P
L
L
S
Q
F
K
V
Q
I
D
Site 86
S1064
Q
F
K
V
Q
I
D
S
Y
E
T
L
Y
E
E
Site 87
Y1065
F
K
V
Q
I
D
S
Y
E
T
L
Y
E
E
V
Site 88
Y1069
I
D
S
Y
E
T
L
Y
E
E
V
C
R
L
E
Site 89
S1095
D
I
R
P
F
K
A
S
L
L
N
I
I
K
R
Site 90
T1116
Q
H
L
V
D
H
V
T
H
S
L
A
N
L
D
Site 91
S1129
L
D
A
F
I
K
K
S
E
S
G
L
L
K
K
Site 92
S1160
M
A
V
K
E
R
Q
S
N
T
D
E
M
F
E
Site 93
T1178
Q
T
I
E
L
L
K
T
Y
E
Q
E
L
P
E
Site 94
T1186
Y
E
Q
E
L
P
E
T
V
F
K
Q
L
E
E
Site 95
T1228
T
L
L
R
Q
R
C
T
A
F
D
A
E
Q
Q
Site 96
S1251
E
A
P
F
R
F
D
S
I
H
P
H
Q
M
L
Site 97
S1270
I
E
I
Q
Q
M
E
S
T
M
A
S
I
S
E
Site 98
S1274
Q
M
E
S
T
M
A
S
I
S
E
S
A
S
L
Site 99
S1276
E
S
T
M
A
S
I
S
E
S
A
S
L
F
E
Site 100
Y1289
F
E
V
N
V
P
D
Y
K
Q
L
R
Q
C
R
Site 101
T1308
Q
L
K
E
L
W
D
T
I
G
M
V
T
S
S
Site 102
S1396
L
M
Q
A
T
G
V
S
F
T
M
D
Q
D
T
Site 103
T1403
S
F
T
M
D
Q
D
T
T
L
A
H
L
L
Q
Site 104
Y1416
L
Q
L
Q
L
H
H
Y
E
D
E
V
R
G
I
Site 105
T1443
T
L
K
E
L
Q
T
T
W
A
G
M
E
F
Q
Site 106
T1457
Q
Y
E
P
H
P
R
T
N
V
P
L
L
C
S
Site 107
S1503
S
G
W
Q
K
K
L
S
T
V
D
A
V
I
S
Site 108
T1520
F
E
V
Q
R
T
W
T
H
L
E
S
I
F
T
Site 109
S1524
R
T
W
T
H
L
E
S
I
F
T
G
S
E
D
Site 110
T1527
T
H
L
E
S
I
F
T
G
S
E
D
I
R
A
Site 111
S1540
R
A
Q
L
P
Q
D
S
K
R
F
E
G
I
D
Site 112
Y1573
T
T
N
K
P
G
L
Y
E
K
L
E
D
I
Q
Site 113
T1596
A
L
A
E
Y
L
D
T
K
R
L
A
F
P
R
Site 114
S1617
S
D
L
L
D
I
L
S
N
G
T
A
P
Q
Q
Site 115
S1630
Q
Q
V
Q
R
H
L
S
K
L
F
D
N
M
A
Site 116
S1646
M
R
F
Q
L
D
A
S
G
E
P
T
K
T
S
Site 117
T1650
L
D
A
S
G
E
P
T
K
T
S
L
G
M
Y
Site 118
T1652
A
S
G
E
P
T
K
T
S
L
G
M
Y
S
K
Site 119
S1653
S
G
E
P
T
K
T
S
L
G
M
Y
S
K
E
Site 120
Y1657
T
K
T
S
L
G
M
Y
S
K
E
E
E
Y
V
Site 121
S1658
K
T
S
L
G
M
Y
S
K
E
E
E
Y
V
A
Site 122
Y1663
M
Y
S
K
E
E
E
Y
V
A
F
S
E
P
C
Site 123
S1667
E
E
E
Y
V
A
F
S
E
P
C
D
C
S
G
Site 124
T1690
V
L
G
H
M
K
A
T
V
R
H
E
M
T
E
Site 125
Y1742
F
A
R
L
E
E
G
Y
E
S
A
M
K
D
Y
Site 126
S1744
R
L
E
E
G
Y
E
S
A
M
K
D
Y
Y
K
Site 127
Y1749
Y
E
S
A
M
K
D
Y
Y
K
K
Q
V
A
Q
Site 128
Y1750
E
S
A
M
K
D
Y
Y
K
K
Q
V
A
Q
L
Site 129
T1759
K
Q
V
A
Q
L
K
T
L
I
T
M
L
I
G
Site 130
S1769
T
M
L
I
G
Q
L
S
K
G
D
R
Q
K
I
Site 131
Y1832
I
C
D
A
Q
F
L
Y
S
Y
E
Y
L
G
N
Site 132
Y1834
D
A
Q
F
L
Y
S
Y
E
Y
L
G
N
T
P
Site 133
Y1836
Q
F
L
Y
S
Y
E
Y
L
G
N
T
P
R
L
Site 134
T1840
S
Y
E
Y
L
G
N
T
P
R
L
V
I
T
P
Site 135
T1846
N
T
P
R
L
V
I
T
P
L
T
D
R
C
Y
Site 136
Y1853
T
P
L
T
D
R
C
Y
I
T
L
T
Q
S
L
Site 137
S1865
Q
S
L
H
L
T
M
S
G
A
P
A
G
P
A
Site 138
Y1902
N
C
S
E
Q
M
D
Y
K
S
C
G
N
I
Y
Site 139
S1954
R
D
K
K
Q
W
F
S
F
L
G
E
E
I
S
Site 140
Y1976
F
I
T
M
N
P
G
Y
A
G
R
T
E
L
P
Site 141
S1988
E
L
P
E
N
L
K
S
L
F
R
P
C
A
M
Site 142
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K
E
N
P
S
K
W
Site 294
S4473
L
K
T
K
E
N
P
S
K
W
V
L
A
G
V
Legend
Confirmed in mammals
Confirmed in related proteins or other species
Predicted by Kinexus P-Site Prediction algorithm
No data/link available
Link available
Products available
2019 Kinexus Bioinformatics Corporation