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Updated November 2019
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Protein Info
Short Name:
STARD9
Full Name:
StAR-related lipid transfer protein 9
Alias:
START domain-containing protein 9
Type:
Mass (Da):
506731
Number AA:
4614
UniProt ID:
Q9P2P6
International Prot ID:
IPI00514002
Sequence:
Retrieve full protein sequence
GO Terms
External Links
Internal Links
Cellular Component:
GO:0005875
GO:0032991
GO:0043234
Uniprot
OncoNet
Molecular Function:
GO:0000166
GO:0003774
GO:0003777
PhosphoSite+
KinaseNET
Biological Process:
GO:0006810
GO:0006996
GO:0007010
Phosida
TranscriptoNet
STRING
Kinexus Products
Info Box
A variety of parameters were considered in the selection of putative P-sites. Confirmed P-Sites have lower Hydrophobicity Scores. The P-site Similarity Score is lower the more that it resembles typical confirmed corresponding P-Ser, P-Thr or P-Tyr sites. The Maximum KInase Score provides the calculated score for highest match of 500 human protein kinases for the amino acid sequence surrounding the target P-site as determined with Kinase Substrate Predictor V2. The Sum KInase Score provides the additive sum of the positive individual Kinase Substrate Predictor V2 scores from 500 human protein kinases. The Conservation Score is the average of the percent similarity of the human P-site with the equivalent P-site in 20 other diverse species. Click the coloured buttons below to retrieve this and other information in the Info Box, or click the orange buttons for relevant links.
Phosphosites
-7
-6
-5
-4
-3
-2
-1
0
1
2
3
4
5
6
7
Expt. conf.
Effect
Kinase
PPase
Kinexus Products
Ref.
Evol.
Kinase Pred.
P-site Match
Site 1
S14
A
V
R
V
R
P
L
S
K
R
E
T
K
E
G
Site 2
T18
R
P
L
S
K
R
E
T
K
E
G
G
R
I
I
Site 3
S50
R
P
D
G
F
G
D
S
R
E
K
V
M
A
F
Site 4
Y73
V
N
P
E
D
P
Q
Y
A
S
Q
D
V
V
F
Site 5
S75
P
E
D
P
Q
Y
A
S
Q
D
V
V
F
Q
D
Site 6
Y102
Y
N
I
C
L
F
A
Y
G
Q
T
G
S
G
K
Site 7
Y111
Q
T
G
S
G
K
T
Y
T
M
L
G
T
P
V
Site 8
T112
T
G
S
G
K
T
Y
T
M
L
G
T
P
V
S
Site 9
S139
V
R
E
K
D
C
A
S
L
P
S
S
C
R
I
Site 10
S142
K
D
C
A
S
L
P
S
S
C
R
I
K
V
S
Site 11
S149
S
S
C
R
I
K
V
S
F
L
E
I
Y
N
E
Site 12
Y154
K
V
S
F
L
E
I
Y
N
E
R
V
R
D
L
Site 13
S165
V
R
D
L
L
K
Q
S
G
Q
K
K
S
Y
T
Site 14
S170
K
Q
S
G
Q
K
K
S
Y
T
L
R
V
R
E
Site 15
T172
S
G
Q
K
K
S
Y
T
L
R
V
R
E
H
P
Site 16
Y184
E
H
P
E
M
G
P
Y
V
Q
G
L
S
Q
H
Site 17
S189
G
P
Y
V
Q
G
L
S
Q
H
V
V
T
N
Y
Site 18
Y196
S
Q
H
V
V
T
N
Y
K
Q
V
I
Q
L
L
Site 19
T212
E
G
I
A
N
R
I
T
A
A
T
H
V
H
E
Site 20
T215
A
N
R
I
T
A
A
T
H
V
H
E
A
S
S
Site 21
S221
A
T
H
V
H
E
A
S
S
R
S
H
A
I
F
Site 22
S222
T
H
V
H
E
A
S
S
R
S
H
A
I
F
T
Site 23
S224
V
H
E
A
S
S
R
S
H
A
I
F
T
I
H
Site 24
S257
N
L
V
D
L
A
G
S
E
R
A
D
P
S
Y
Site 25
S263
G
S
E
R
A
D
P
S
Y
C
K
D
R
I
A
Site 26
Y264
S
E
R
A
D
P
S
Y
C
K
D
R
I
A
E
Site 27
S293
I
S
T
L
A
Q
N
S
Q
V
F
S
S
C
Q
Site 28
S297
A
Q
N
S
Q
V
F
S
S
C
Q
S
L
N
S
Site 29
S298
Q
N
S
Q
V
F
S
S
C
Q
S
L
N
S
S
Site 30
S301
Q
V
F
S
S
C
Q
S
L
N
S
S
V
S
N
Site 31
S304
S
S
C
Q
S
L
N
S
S
V
S
N
G
G
D
Site 32
S305
S
C
Q
S
L
N
S
S
V
S
N
G
G
D
S
Site 33
S307
Q
S
L
N
S
S
V
S
N
G
G
D
S
G
I
Site 34
S312
S
V
S
N
G
G
D
S
G
I
L
S
S
P
S
Site 35
S316
G
G
D
S
G
I
L
S
S
P
S
G
T
S
S
Site 36
S317
G
D
S
G
I
L
S
S
P
S
G
T
S
S
G
Site 37
S319
S
G
I
L
S
S
P
S
G
T
S
S
G
G
A
Site 38
T321
I
L
S
S
P
S
G
T
S
S
G
G
A
P
S
Site 39
S323
S
S
P
S
G
T
S
S
G
G
A
P
S
R
R
Site 40
S328
T
S
S
G
G
A
P
S
R
R
Q
S
Y
I
P
Site 41
S332
G
A
P
S
R
R
Q
S
Y
I
P
Y
R
D
S
Site 42
Y333
A
P
S
R
R
Q
S
Y
I
P
Y
R
D
S
V
Site 43
Y336
R
R
Q
S
Y
I
P
Y
R
D
S
V
L
T
W
Site 44
S339
S
Y
I
P
Y
R
D
S
V
L
T
W
L
L
K
Site 45
T342
P
Y
R
D
S
V
L
T
W
L
L
K
D
S
L
Site 46
S348
L
T
W
L
L
K
D
S
L
G
G
N
S
K
T
Site 47
S366
A
I
T
R
Q
T
M
S
T
L
R
Y
A
S
S
Site 48
T367
I
T
R
Q
T
M
S
T
L
R
Y
A
S
S
A
Site 49
Y370
Q
T
M
S
T
L
R
Y
A
S
S
A
K
N
I
Site 50
S372
M
S
T
L
R
Y
A
S
S
A
K
N
I
I
N
Site 51
S373
S
T
L
R
Y
A
S
S
A
K
N
I
I
N
K
Site 52
S395
P
F
Q
K
N
F
S
S
L
S
D
E
N
L
K
Site 53
S397
Q
K
N
F
S
S
L
S
D
E
N
L
K
E
L
Site 54
T416
E
L
K
V
D
Q
L
T
K
D
W
T
Q
K
W
Site 55
S434
Q
A
L
M
E
H
Y
S
V
D
I
N
R
R
R
Site 56
T500
P
A
R
G
A
R
C
T
V
N
G
R
E
V
T
Site 57
T507
T
V
N
G
R
E
V
T
A
S
C
R
L
T
Q
Site 58
S509
N
G
R
E
V
T
A
S
C
R
L
T
Q
G
A
Site 59
T513
V
T
A
S
C
R
L
T
Q
G
A
V
I
T
L
Site 60
S552
E
A
A
A
G
R
G
S
L
E
W
L
D
L
D
Site 61
S570
A
A
S
R
L
G
L
S
P
L
L
W
K
E
R
Site 62
T590
Q
C
D
E
D
H
Q
T
P
R
D
G
E
T
S
Site 63
S597
T
P
R
D
G
E
T
S
H
R
A
Q
I
Q
Q
Site 64
Y608
Q
I
Q
Q
Q
Q
S
Y
V
E
D
L
R
H
Q
Site 65
T650
Q
C
L
L
R
E
E
T
W
L
A
S
L
Q
Q
Site 66
S654
R
E
E
T
W
L
A
S
L
Q
Q
Q
Q
Q
E
Site 67
S688
T
D
P
E
I
Q
P
S
P
F
V
Q
S
Q
K
Site 68
S693
Q
P
S
P
F
V
Q
S
Q
K
R
V
V
H
L
Site 69
T707
L
Q
L
L
R
R
H
T
L
R
A
A
E
R
N
Site 70
S721
N
V
R
R
K
K
V
S
F
Q
L
E
R
I
I
Site 71
T744
Q
K
R
L
E
K
L
T
T
L
C
W
L
Q
D
Site 72
T745
K
R
L
E
K
L
T
T
L
C
W
L
Q
D
D
Site 73
T754
C
W
L
Q
D
D
S
T
Q
E
P
P
Y
Q
V
Site 74
Y759
D
S
T
Q
E
P
P
Y
Q
V
L
S
P
D
A
Site 75
S763
E
P
P
Y
Q
V
L
S
P
D
A
T
V
P
R
Site 76
T767
Q
V
L
S
P
D
A
T
V
P
R
P
P
C
R
Site 77
S775
V
P
R
P
P
C
R
S
K
L
T
S
C
S
S
Site 78
T778
P
P
C
R
S
K
L
T
S
C
S
S
L
S
P
Site 79
S779
P
C
R
S
K
L
T
S
C
S
S
L
S
P
Q
Site 80
S781
R
S
K
L
T
S
C
S
S
L
S
P
Q
R
L
Site 81
S782
S
K
L
T
S
C
S
S
L
S
P
Q
R
L
C
Site 82
S784
L
T
S
C
S
S
L
S
P
Q
R
L
C
S
K
Site 83
S790
L
S
P
Q
R
L
C
S
K
H
M
P
Q
L
H
Site 84
S802
Q
L
H
S
I
F
L
S
W
D
P
S
T
T
L
Site 85
S806
I
F
L
S
W
D
P
S
T
T
L
P
P
R
P
Site 86
T807
F
L
S
W
D
P
S
T
T
L
P
P
R
P
D
Site 87
T808
L
S
W
D
P
S
T
T
L
P
P
R
P
D
P
Site 88
T816
L
P
P
R
P
D
P
T
H
Q
T
S
E
K
T
Site 89
S820
P
D
P
T
H
Q
T
S
E
K
T
S
S
E
E
Site 90
S824
H
Q
T
S
E
K
T
S
S
E
E
H
L
P
Q
Site 91
S825
Q
T
S
E
K
T
S
S
E
E
H
L
P
Q
A
Site 92
S834
E
H
L
P
Q
A
A
S
Y
P
A
R
T
G
C
Site 93
Y835
H
L
P
Q
A
A
S
Y
P
A
R
T
G
C
L
Site 94
S850
R
K
N
G
L
H
S
S
G
H
G
Q
P
C
T
Site 95
T857
S
G
H
G
Q
P
C
T
A
R
A
A
L
A
R
Site 96
T875
S
A
P
D
A
C
L
T
M
S
P
N
S
V
G
Site 97
S880
C
L
T
M
S
P
N
S
V
G
I
Q
E
M
E
Site 98
S898
K
Q
P
H
Q
M
V
S
Q
G
L
A
S
L
R
Site 99
S903
M
V
S
Q
G
L
A
S
L
R
K
S
A
N
K
Site 100
S907
G
L
A
S
L
R
K
S
A
N
K
L
K
P
R
Site 101
T924
P
K
I
F
T
S
T
T
Q
T
R
G
A
K
G
Site 102
T926
I
F
T
S
T
T
Q
T
R
G
A
K
G
L
A
Site 103
S936
A
K
G
L
A
D
P
S
H
T
Q
A
G
W
R
Site 104
S961
A
K
G
A
S
C
N
S
L
Y
P
H
G
P
R
Site 105
Y963
G
A
S
C
N
S
L
Y
P
H
G
P
R
Q
T
Site 106
T970
Y
P
H
G
P
R
Q
T
A
G
H
G
K
A
V
Site 107
T979
G
H
G
K
A
V
K
T
F
W
T
E
Y
K
P
Site 108
T982
K
A
V
K
T
F
W
T
E
Y
K
P
P
S
P
Site 109
Y984
V
K
T
F
W
T
E
Y
K
P
P
S
P
S
R
Site 110
S988
W
T
E
Y
K
P
P
S
P
S
R
A
S
K
R
Site 111
S990
E
Y
K
P
P
S
P
S
R
A
S
K
R
H
Q
Site 112
S993
P
P
S
P
S
R
A
S
K
R
H
Q
R
V
L
Site 113
T1008
A
T
R
V
R
N
I
T
K
K
S
S
H
L
P
Site 114
S1011
V
R
N
I
T
K
K
S
S
H
L
P
L
G
S
Site 115
S1012
R
N
I
T
K
K
S
S
H
L
P
L
G
S
P
Site 116
S1018
S
S
H
L
P
L
G
S
P
L
K
R
Q
Q
N
Site 117
T1031
Q
N
T
R
D
P
D
T
M
V
P
L
T
D
F
Site 118
T1036
P
D
T
M
V
P
L
T
D
F
S
P
V
M
D
Site 119
S1039
M
V
P
L
T
D
F
S
P
V
M
D
H
S
R
Site 120
S1045
F
S
P
V
M
D
H
S
R
E
K
D
N
D
L
Site 121
S1053
R
E
K
D
N
D
L
S
D
T
D
S
N
Y
S
Site 122
T1055
K
D
N
D
L
S
D
T
D
S
N
Y
S
L
D
Site 123
S1057
N
D
L
S
D
T
D
S
N
Y
S
L
D
S
L
Site 124
Y1059
L
S
D
T
D
S
N
Y
S
L
D
S
L
S
C
Site 125
S1060
S
D
T
D
S
N
Y
S
L
D
S
L
S
C
V
Site 126
S1063
D
S
N
Y
S
L
D
S
L
S
C
V
Y
A
K
Site 127
S1091
D
F
P
E
P
E
N
S
E
S
D
D
S
Q
L
Site 128
S1093
P
E
P
E
N
S
E
S
D
D
S
Q
L
S
E
Site 129
S1096
E
N
S
E
S
D
D
S
Q
L
S
E
D
S
L
Site 130
S1099
E
S
D
D
S
Q
L
S
E
D
S
L
A
E
K
Site 131
S1102
D
S
Q
L
S
E
D
S
L
A
E
K
R
Y
Q
Site 132
Y1108
D
S
L
A
E
K
R
Y
Q
S
P
K
N
R
L
Site 133
S1110
L
A
E
K
R
Y
Q
S
P
K
N
R
L
G
G
Site 134
T1121
R
L
G
G
N
R
P
T
N
N
R
G
Q
P
R
Site 135
T1129
N
N
R
G
Q
P
R
T
R
T
R
A
S
V
R
Site 136
T1131
R
G
Q
P
R
T
R
T
R
A
S
V
R
G
F
Site 137
S1134
P
R
T
R
T
R
A
S
V
R
G
F
T
A
A
Site 138
T1139
R
A
S
V
R
G
F
T
A
A
S
D
S
D
L
Site 139
S1144
G
F
T
A
A
S
D
S
D
L
L
A
Q
T
H
Site 140
T1150
D
S
D
L
L
A
Q
T
H
R
S
F
S
L
D
Site 141
S1155
A
Q
T
H
R
S
F
S
L
D
S
L
I
D
A
Site 142
S1158
H
R
S
F
S
L
D
S
L
I
D
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T3337
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S3396
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S3427
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S3441
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T3551
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T3558
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S3611
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S3613
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T3624
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T3645
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S3650
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T3652
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T3655
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T3658
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S3664
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T3682
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T3684
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S3686
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T3719
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S3725
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S3727
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S3731
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S3746
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S3765
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S3774
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S3776
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S3777
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S3780
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S3786
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T3787
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S3788
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S3794
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T3817
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S3819
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S3828
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S3831
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T3833
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S3835
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T3839
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T3860
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S3979
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S4000
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S4292
R
I
H
Q
K
I
I
S
Q
L
L
K
E
E
D
Site 598
T4303
K
E
E
D
K
L
H
T
L
A
N
S
S
S
L
Site 599
S4307
K
L
H
T
L
A
N
S
S
S
L
C
T
S
S
Site 600
S4308
L
H
T
L
A
N
S
S
S
L
C
T
S
S
N
Site 601
S4309
H
T
L
A
N
S
S
S
L
C
T
S
S
N
G
Site 602
T4312
A
N
S
S
S
L
C
T
S
S
N
G
S
L
S
Site 603
S4313
N
S
S
S
L
C
T
S
S
N
G
S
L
S
S
Site 604
S4314
S
S
S
L
C
T
S
S
N
G
S
L
S
S
G
Site 605
S4317
L
C
T
S
S
N
G
S
L
S
S
G
M
T
S
Site 606
S4319
T
S
S
N
G
S
L
S
S
G
M
T
S
G
Y
Site 607
S4320
S
S
N
G
S
L
S
S
G
M
T
S
G
Y
N
Site 608
T4323
G
S
L
S
S
G
M
T
S
G
Y
N
S
S
P
Site 609
S4324
S
L
S
S
G
M
T
S
G
Y
N
S
S
P
A
Site 610
Y4326
S
S
G
M
T
S
G
Y
N
S
S
P
A
L
S
Site 611
S4328
G
M
T
S
G
Y
N
S
S
P
A
L
S
G
Q
Site 612
S4329
M
T
S
G
Y
N
S
S
P
A
L
S
G
Q
L
Site 613
S4333
Y
N
S
S
P
A
L
S
G
Q
L
Q
F
P
E
Site 614
S4350
G
H
T
N
L
P
D
S
R
D
V
W
I
G
D
Site 615
S4363
G
D
E
R
G
G
H
S
A
V
R
K
N
S
A
Site 616
S4369
H
S
A
V
R
K
N
S
A
Y
S
H
R
A
S
Site 617
Y4371
A
V
R
K
N
S
A
Y
S
H
R
A
S
L
G
Site 618
S4372
V
R
K
N
S
A
Y
S
H
R
A
S
L
G
S
Site 619
S4376
S
A
Y
S
H
R
A
S
L
G
S
C
C
C
S
Site 620
S4379
S
H
R
A
S
L
G
S
C
C
C
S
P
S
S
Site 621
T4392
S
S
L
S
S
L
G
T
C
F
S
S
S
Y
Q
Site 622
S4395
S
S
L
G
T
C
F
S
S
S
Y
Q
D
L
A
Site 623
S4396
S
L
G
T
C
F
S
S
S
Y
Q
D
L
A
K
Site 624
S4397
L
G
T
C
F
S
S
S
Y
Q
D
L
A
K
H
Site 625
Y4445
E
E
Q
A
V
Q
L
Y
Y
K
V
F
S
P
T
Site 626
Y4446
E
Q
A
V
Q
L
Y
Y
K
V
F
S
P
T
R
Site 627
S4450
Q
L
Y
Y
K
V
F
S
P
T
R
H
G
F
L
Site 628
Y4482
D
P
T
V
W
P
L
Y
Y
K
P
I
Q
T
A
Site 629
Y4483
P
T
V
W
P
L
Y
Y
K
P
I
Q
T
A
R
Site 630
T4496
A
R
L
H
Q
R
V
T
N
S
I
S
L
V
Y
Site 631
S4532
E
A
K
E
G
H
L
S
V
M
A
A
Q
S
V
Site 632
T4542
A
A
Q
S
V
Y
D
T
S
M
P
R
P
S
R
Site 633
S4543
A
Q
S
V
Y
D
T
S
M
P
R
P
S
R
K
Site 634
S4548
D
T
S
M
P
R
P
S
R
K
M
V
R
G
E
Site 635
S4595
G
F
P
P
Q
L
L
S
S
F
I
K
R
Q
P
Site 636
S4596
F
P
P
Q
L
L
S
S
F
I
K
R
Q
P
L
Legend
Confirmed in mammals
Confirmed in related proteins or other species
Predicted by Kinexus P-Site Prediction algorithm
No data/link available
Link available
Products available
2019 Kinexus Bioinformatics Corporation