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Search by protein name, UniProt number, IPI number, or 15 AA P-site sequence.
Updated November 2019
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Protein Info
Short Name:
USP21
Full Name:
Ubiquitin carboxyl-terminal hydrolase 21
Alias:
Deubiquitinating enzyme 21; Ubiquitin specific peptidase 21; Ubiquitin thioesterase 21; Ubiquitin-specific-processing protease 21; USP16; USP23
Type:
EC 3.1.2.15; Protease; Ubiquitin conjugating system
Mass (Da):
62637
Number AA:
565
UniProt ID:
Q9UK80
International Prot ID:
IPI00007262
Sequence:
Retrieve full protein sequence
GO Terms
External Links
Internal Links
Cellular Component:
GO:0005634
Uniprot
OncoNet
Molecular Function:
GO:0019784
GO:0003713
GO:0004221
PhosphoSite+
KinaseNET
Biological Process:
GO:0016578
GO:0045893
GO:0006350
Phosida
TranscriptoNet
STRING
Kinexus Products
Info Box
A variety of parameters were considered in the selection of putative P-sites. Confirmed P-Sites have lower Hydrophobicity Scores. The P-site Similarity Score is lower the more that it resembles typical confirmed corresponding P-Ser, P-Thr or P-Tyr sites. The Maximum KInase Score provides the calculated score for highest match of 500 human protein kinases for the amino acid sequence surrounding the target P-site as determined with Kinase Substrate Predictor V2. The Sum KInase Score provides the additive sum of the positive individual Kinase Substrate Predictor V2 scores from 500 human protein kinases. The Conservation Score is the average of the percent similarity of the human P-site with the equivalent P-site in 20 other diverse species. Click the coloured buttons below to retrieve this and other information in the Info Box, or click the orange buttons for relevant links.
Phosphosites
-7
-6
-5
-4
-3
-2
-1
0
1
2
3
4
5
6
7
Expt. conf.
Effect
Kinase
PPase
Kinexus Products
Ref.
Evol.
Kinase Pred.
P-site Match
Site 1
S25
N
I
Q
P
R
V
G
S
K
L
P
F
A
P
R
Site 2
S35
P
F
A
P
R
A
R
S
K
E
R
R
N
P
A
Site 3
S43
K
E
R
R
N
P
A
S
G
P
N
P
M
L
R
Site 4
T73
L
E
L
G
R
G
R
T
S
G
P
R
P
R
G
Site 5
S74
E
L
G
R
G
R
T
S
G
P
R
P
R
G
P
Site 6
S93
H
G
V
P
L
P
G
S
P
P
P
T
V
A
L
Site 7
T106
A
L
P
L
P
S
R
T
N
L
A
R
S
K
S
Site 8
S111
S
R
T
N
L
A
R
S
K
S
V
S
S
G
D
Site 9
S113
T
N
L
A
R
S
K
S
V
S
S
G
D
L
R
Site 10
S115
L
A
R
S
K
S
V
S
S
G
D
L
R
P
M
Site 11
S116
A
R
S
K
S
V
S
S
G
D
L
R
P
M
G
Site 12
S140
G
E
L
G
A
A
L
S
R
L
A
L
R
P
E
Site 13
T150
A
L
R
P
E
P
P
T
L
R
R
S
T
S
L
Site 14
S154
E
P
P
T
L
R
R
S
T
S
L
R
R
L
G
Site 15
T155
P
P
T
L
R
R
S
T
S
L
R
R
L
G
G
Site 16
S156
P
T
L
R
R
S
T
S
L
R
R
L
G
G
F
Site 17
T168
G
G
F
P
G
P
P
T
L
F
S
I
R
T
E
Site 18
S171
P
G
P
P
T
L
F
S
I
R
T
E
P
P
A
Site 19
T174
P
T
L
F
S
I
R
T
E
P
P
A
S
H
G
Site 20
S179
I
R
T
E
P
P
A
S
H
G
S
F
H
M
I
Site 21
S182
E
P
P
A
S
H
G
S
F
H
M
I
S
A
R
Site 22
S187
H
G
S
F
H
M
I
S
A
R
S
S
E
P
F
Site 23
S190
F
H
M
I
S
A
R
S
S
E
P
F
Y
S
D
Site 24
S191
H
M
I
S
A
R
S
S
E
P
F
Y
S
D
D
Site 25
Y195
A
R
S
S
E
P
F
Y
S
D
D
K
M
A
H
Site 26
S196
R
S
S
E
P
F
Y
S
D
D
K
M
A
H
H
Site 27
T204
D
D
K
M
A
H
H
T
L
L
L
G
S
G
H
Site 28
S231
N
A
V
L
Q
C
L
S
S
T
R
P
L
R
D
Site 29
S274
G
A
L
W
H
P
D
S
C
E
A
V
N
P
T
Site 30
Y290
F
R
A
V
F
Q
K
Y
V
P
S
F
S
G
Y
Site 31
S293
V
F
Q
K
Y
V
P
S
F
S
G
Y
S
Q
Q
Site 32
Y297
Y
V
P
S
F
S
G
Y
S
Q
Q
D
A
Q
E
Site 33
S335
L
A
N
G
P
V
P
S
P
P
R
R
G
G
A
Site 34
S350
L
L
E
E
P
E
L
S
D
D
D
R
A
N
L
Site 35
Y362
A
N
L
M
W
K
R
Y
L
E
R
E
D
S
K
Site 36
S368
R
Y
L
E
R
E
D
S
K
I
V
D
L
F
V
Site 37
S431
T
K
E
E
E
L
E
S
E
N
A
P
V
C
D
Site 38
T444
C
D
R
C
R
Q
K
T
R
S
T
K
K
L
T
Site 39
S446
R
C
R
Q
K
T
R
S
T
K
K
L
T
V
Q
Site 40
T447
C
R
Q
K
T
R
S
T
K
K
L
T
V
Q
R
Site 41
T451
T
R
S
T
K
K
L
T
V
Q
R
F
P
R
I
Site 42
S467
V
L
H
L
N
R
F
S
A
S
R
G
S
I
K
Site 43
S472
R
F
S
A
S
R
G
S
I
K
K
S
S
V
G
Site 44
S476
S
R
G
S
I
K
K
S
S
V
G
V
D
F
P
Site 45
S477
R
G
S
I
K
K
S
S
V
G
V
D
F
P
L
Site 46
S488
D
F
P
L
Q
R
L
S
L
G
D
F
A
S
D
Site 47
S494
L
S
L
G
D
F
A
S
D
K
A
G
S
P
V
Site 48
S499
F
A
S
D
K
A
G
S
P
V
Y
Q
L
Y
A
Site 49
Y502
D
K
A
G
S
P
V
Y
Q
L
Y
A
L
C
N
Site 50
Y505
G
S
P
V
Y
Q
L
Y
A
L
C
N
H
S
G
Site 51
S535
G
W
H
V
Y
N
D
S
R
V
S
P
V
S
E
Site 52
S538
V
Y
N
D
S
R
V
S
P
V
S
E
N
Q
V
Site 53
S541
D
S
R
V
S
P
V
S
E
N
Q
V
A
S
S
Site 54
S548
S
E
N
Q
V
A
S
S
E
G
Y
V
L
F
Y
Site 55
Y555
S
E
G
Y
V
L
F
Y
Q
L
M
Q
E
P
P
Legend
Confirmed in mammals
Confirmed in related proteins or other species
Predicted by Kinexus P-Site Prediction algorithm
No data/link available
Link available
Products available
2019 Kinexus Bioinformatics Corporation